Valid Names Results
Trionymus perrisii (Signoret, 1875) (Pseudococcidae: Trionymus)This catalog page aggregates the information for all subspecies of Trionymus perrisii. If you are looking for information specific to the binomial, click on the link to the nominal trinomial below.
Subspecies:
Nomenclatural History
- Westwoodia perrisii Signoret 1875c: 337. Type data: FRANCE: Mont-de-Marsan, on Calamagrostis arundinacea. Lectotype, female, by subsequent designation (BenDovMa1995,254). Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; accepted valid name Illustr.
- Bergrothia perrisii (Signoret, 1875); Kraatz 1888a: 360. change of combination
- Signoretia perrisii (Signoret, 1875); Kraatz 1888: 176. change of combination
- Dactylopius hibernicus Newstead 1895: 167. Type data: IRELAND: Antrim, Ballintoy, on grass. Syntypes, female, Type depository: London: The Natural History Museum, England, UK; junior synonym (discovered by Willia1962, 63).
- Dactylopius perrisii (Signoret, 1875); Cockerell 1897: 201. change of combination
- Bergrothiella perrisii (Signoret, 1875); Reitter 1898: 54. change of combination
- Trionymus perrisii (Signoret, 1875); Berg 1899: 78. change of combination
- Pseudococcus hibernicus (Newstead, 1895); Fernald 1903b: 103. change of combination
- Pseudococcus pulverarius; Green 1915a: 178. misidentification (discovered by Willia1962, 63).
- Pseudococcus (Trionymus) tomlini Green 1925b: 525. Type data: CHANNEL ISLANDS: Guernsey, Port Grat, on Psamma (Ammophila) arenaria. Syntypes, female, by subsequent designation Type depository: London: The Natural History Museum, England, UK; junior synonym (discovered by DanzigGa2015, 164). Illustr.
- Pseudococcus (Trionymus) californicus; Green 1925b: 522. misidentification
- Trionymus hibernicus (Newstead, 1895); Green 1926: 183. change of combination
- Bergrothura perrisii (Signoret, 1875); Strand 1928: 47. change of combination
- Trionymus perrisi (Signoret, 1875); Goux 1931: 4. misspelling of species epithet
- Erium perrisii (Signoret, 1875); Lindinger 1935a: 122. change of combination
- Trionymus internodii; Kiritchenko 1940: 123. misidentification (discovered by Borchs1949, 145).
- Trionymus perrisii bassensis Goux 1941a: 38. Type data: FRANCE: Bassens, Savoie, on grasses. Syntypes, female, Type depository: Paris: Museum National d'Histoire naturelle, France; accepted valid name Illustr.
- Trionymus perrisii perrisii (Signoret, 1875); Goux 1941: 38. change in status (level)
- Trionymus perrisi bassensis Goux, 1941; Goux 1941: 38. misspelling of species epithet
- Pseudococcus perrisii (Signoret, 1875); Borchsenius 1949: 142. change of combination
- Pseudococcus calamagrostidis Borchsenius 1949: 143. Type data: KYRGYZSTAN: Bank of Kugartka river, 35 km of Dzhalal Abad, on Calamagrostis. Lectotype, female, by subsequent designation (Danzig1997a,98). Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; junior synonym (discovered by DanzigGa2015, 164).
- Pseudococcus graminellus Borchsenius 1949: 145. Type data: RUSSIA: Leningrad] Oblast, Luga, on leaf sheaths of grass.. Lectotype, female, by subsequent designation (Danzig1997a,101). Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; junior synonym (discovered by Danzig1997a, 101).
- Pseudococcus implicatus Borchsenius 1949: 144. Type data: KAZAKHSTAN: Aktyubinsk District, Alga, on grass.. Lectotype, female, by subsequent designation (Danzig1997a,96). Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; junior synonym (discovered by Gavril2003a, 108).
- Trionymus tomlini (Green, 1925); Williams 1962: 72. change of combination
- Trionymus graminellus (Borchsenius, 1949); Ter-Grigorian 1966: 85. change of combination
- Trionymus calamagrostidis (Borchsenius, 1949); Matesova 1968: 103. change of combination
- Trionymus Implicatus (Borchsenius, 1949); Matesova 1968: 104. change of combination
- Trionymus orientalis Matesova 1968: 105. nomen nudum
- Trionymus orientalis Matesova 1968a: 152. Type data: KAZAKHSTAN: near Zyryanovsk Berezovka River, on Agropyron sp.. Holotype, female, Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; synonym and homonym (discovered by Danzig1997a, 101). Notes: synonym and homonym of Trionymus orientalis (Maskell) Illustr.
- Pseudococcus pseudoperrisii Rasina 1971: 3. Type data: LATVIA: Barta, on Luzula pilosa. Holotype, female, Type depository: Riga: Latvian National Museum of Natural History, Latvia; junior synonym
- Trionymus matesovai Kozár & Walter 1985: 73. replacement name that is a synonym Replacement name for Trionymus orientalis Matesova
- Trionymus perrisi (Signoret, 1875); Kozár & Walter 1986: 115. misspelling of species epithet
- Trionymus matesovae Kozár & Walter, 1985; Danzig 1997a: 101. emendation that is justified
- Trionoymus perrisii; Kozár et al. 2013: 94. misspelling of genus name
Common Names
- Perris' grass mealybug KosztaKo1988F
- Guernsey grass mealybug KosztaKo1988F
Ecological Associates
Hosts:
Families: 7 | Genera: 37
- Apiaceae
- Caucalis | KaydanUlEr2007
- Asteraceae
- Centaurea parlatoris | MazzeoRuLo1997
- Hypochoeris | BenDov1994 KosztaKo1988F
- Leontodon | BenDov1994 Koteja1969a BugajNJuKa2021
- Urospermum picroides | SzitaFeBe2017
- Cyperaceae
- Carex spicata | BenDov1994 Ossian1985 | (= Carex cotigua)
- Euphorbiaceae
- Euphorbia pithyusa | Balach1930 BenDov1994
- Juncaceae
- Luzula pilosa | BenDov1994 Rasina1971
- Poaceae
- Aegilops | KaydanKo2010a
- Agropyron | BenDov1994 KaydanKiKo2005 KaydanUlEr2007 Mateso1968a
- Agropyron cristatum | Danzig1997a
- Agrostis canina | BenDov1994 Kozar1985 | (= Agrostis tenuis)
- Agrostis capillaris | BenDov1994 KotejaZa1983 | (= Agrostis vulgaris)
- Agrostis stolonifera | BenDov1994 KozarWa1986
- Ammophila arenaria | BenDov1994 Green1925b
- Anthoxanthum odoratum | BenDov1994 KotejaZa1983
- Arrhenatherum elatius | BenDov1994 BoratyWi1964a
- Bambusa | BugajNJuKa2021
- Brachypodium | BenDov1994 KosztaKo1988F
- Brachypodium phoenicoides | KozarFr1995
- Brachypodium pinnatum | BugajNJuKa2021
- Bromus | BenDov1994 KosztaKo1988F
- Bromus erectus | BugajNJuKa2021
- Bromus inermis | BugajNJuKa2021
- Calamagrostis | BenDov1994 Borchs1949 Danzig1997a
- Calamagrostis arundinacea | BenDov1994
- Calamagrostis canescens | Gertss2005a
- Calamagrostis epigeios | BenDov1994 Danzig1959
- Calamagrostis varia | BenDov1994 Koteja1969a
- Corynephorus canescens | BenDov1994 Schmut1952
- Cynodon dactylon | BenDov1994 KozarTrPe1984 Marott1987a MazzeoLoRu1994
- Dactylis glomerata | BatsanKaKi2017
- Danthonia decumbens | BenDov1994 KotejaZa1969 KotejaZa1983 | (= Sieglingia decumbens)
- Deschampsia | BugajNJuKa2021
- Deschampsia cespitosa | BenDov1994 Willia1962
- Deschampsia flexuosa | BugajNJuKa2021
- Elymus | KaydanKiKo2005 KaydanUlEr2007
- Elymus caninus | BugajNJuKa2021
- Elymus repens | BenDov1994 Danzig1959 KozarDr1991 KozarGuBa1994 Willia1962 | (= Agropyron repens)(= Elytrigia repens)
- Elymus tsukushiensis | KwonDaPa2003a Suh2020 | (= Agropyron tsukushiense
- Festuca | BenDov1994 KozarGuBa1994 KozarWa1986 TerGri1966
- Festuca ovina | BenDov1994 KotejaZa1969 KotejaZa1983
- Festuca pratensis | BenDov1994 KozarWa1986
- Festuca pseudovina | BenDov1994 KozarTzVi1979
- Festuca rubra | BenDov1994 Willia1962
- Holcus | BenDov1994 Willia1962
- Holcus lanatus | BenDov1994 KotejaZa1983
- Holcus mollis | BenDov1994 BoratyWi1964a
- Hordeum murinum | KaydanKiKo2005 KaydanUlEr2007
- Koeleria | BugajNJuKa2021
- Koeleria glauca | BugajNJuKa2021
- Leymus chinensis | BenDov1994 TangHaTa1992 | (= Aneurolepidium chinense)
- Lolium perenne | BugajNJuKa2021
- Lolium rigidum | SzitaFeBe2017
- Molinia caerulea | BenDov1994 Marcha1908 | (= Festuca caerulea)
- Nardus stricta | BugajNJuKa2021
- Phleum phleoides | BenDov1994 Mateso1968
- Phleum pratense | BugajNJuKa2021
- Phragmites | BenDov1994
- Phragmites australis | BenDov1994 Danzig1997a Mateso1968 | (= Phragmites communis)
- Poa | BenDov1994 TerGri1966
- Poa annua | BenDov1994 Koteja1969
- Poa compressa | BenDov1994 KotejaZa1983
- Poa nemoralis | KozarOrKo1977 BenDov1994 Kozar1985a | (= Agrostis alba),
- Poa pratensis | BugajNJuKa2021
- Poa versicolor | BenDov1994 Danzig1997a Mateso1968 | (= Poa stepposa)
- Poaceae | BenDov1994 Goux1941a BenDov1994 Borchs1949 TerGri1966
- Sphagnaceae
- Sphagnum | BenDov1994 Signor1875
Foes:
Families: 1 | Genera: 5
- Encyrtidae
- Ericydnus robustior | Trjapi1989
- Mahencyrtus comara | Trjapi1989
- Mayridia myrlea | Schmut1955b Trjapi1989 | (= Mayridia bifasciatella)
- Pseudococcobius obenbergeri | Trjapi1989
- Rhopus brachypterus | Schmut1955b Trjapi1989 | (= Xanthoencyrtus brachypterus)
- Rhopus parvulus | Schmut1955b | (= Xanthoencyrtus parvulus)
- Rhopus semiapterus | Schmut1955b | (= Xanthoencyrtus semiapterus)
Geographic Distribution
Countries: 34
- Armenia | BenDov1994 Borchs1949 Danzig1997a TerGri1966
- Austria | BugajNJuKa2021
- Bulgaria | BenDov1994 KozarTzVi1979
- China | BenDov1994 Tang1992 TangHaTa1992
- Czech Republic | BenDov1994 KosztaKo1988F
- Denmark | BenDov1994 KozarzRe1975
- France | BenDov1994 Goux1941a | Balach1930 BenDov1994 Foldi2001 Goux1931
- Georgia (Republic of) | BatsanKaKi2017
- Germany | BenDov1994 Schmut1952 Schmut1955b Schmut1980
- Greece | SzitaFeBe2017
- Hungary | BenDov1994 KozarKoSc1999 KozarOrKo1977 KozarWa1986
- Ireland | BenDov1994 Willia1962
- Italy | BenDov1994 KozarTrPe1984 LongoMaPe1995 Marott1987a MazzeoLoRu1994
- Sicily | MazzeoRuLo1997 RussoMa1997
- Kazakhstan | BenDov1994 Borchs1949 Danzig1997a Mateso1968
- Kyrgyzstan (=Kirgizia) | BenDov1994 Borchs1949
- Latvia | BenDov1994 Rasina1971
- Mongolia | Danzig1997a | BenDov1994 Danzig1975 Danzig1980b
- Morocco | Gavril2016
- Netherlands | BenDov1994 Reyne1951 Willia1962
- North Korea | BenDov1994 Danzig1980b
- Poland | BenDov1994 Borchs1949 GolanLaJa2001 Koteja1974b KotejaZa1969 KotejaZa1979 Lagows1996
- Portugal | FrancoRuMa2011 KozarFr1995
- Romania | BenDov1994 Kozar1985
- Russia
- Irkutsk Oblast | Danzig1997a
- Kaliningrad Oblast | Danzig1997a
- Krasnoyarsk Kray | Danzig1997a
- Kuril Islands | BenDov1994 Danzig1980b
- Omsk Oblast | Danzig1997a
- Primor'ye Kray | BenDov1994 Danzig1980b
- Sakhalin Oblast | BenDov1994 Danzig1980b
- Sakhalin Oblast | Danzig1997a
- St. Petersburg (=Leningrad) Oblast | BenDov1994 Borchs1949
- Tatarstan (=Tatar) AR | BenDov1994 Kozar1985a
- Tula Oblast | Danzig1997a
- Voronoezh Oblast | Danzig1997a Gavril2003a
- Yakutia-Sakha (=Yakut) AR | Danzig1997a
- Slovakia | BugajNJuKa2021
- Slovenia | Seljak2010
- South Korea | ESKSAE1994 KwonDaPa2003a
- Sweden | Gertss2005a
- Switzerland | KozarGuBa1994
- Tajikistan (=Tadzhikistan) | Danzig1997a
- Turkey | KaydanKiKo2005 KaydanKoYa2001 KaydanUlEr2007
- Ukraine | BenDov1994 Danzig1997a Terezn1975
- Krym (=Crimea) Oblast | Danzig1997a
- United Kingdom
- England | Newste1895
- Scotland | BenDov1994 Willia1962
- Yugoslavia | BenDov1994 Kozar1983a
Keys
- YerlikBaKa2021: pp.326-327 ( Adult (F) ) [Trionymus of the Palearctic region]
- DanzigGa2015: pp.99-106 ( Adult (F) ) [Palaearctic Trionymus species]
- KwonDaPa2003a: pp.415 ( Adult (F) ) [Trionymus species of Korea]
- KwonDaPa2003a: pp.415 ( Adult (F) ) [Trionymus species of Korea] Key as: Trionymus tomlini
- Danzig1997a: pp.96 ( Adult (F) ) [Palaearctic region] Key as: Trionymus calamagrostidis
- Danzig1997a: pp.96 ( Adult (F) ) [Palaearctic region] Key as: Trionymus matesovae
- Danzig1997a: pp.96 ( Adult (F) ) [Palaearctic region]
- Danzig1997a: pp.96 ( Adult (F) ) [Palaearctic region] Key as: Trionymus tomlini
- Tang1992: pp.151 ( Adult (F) ) [Palaearctic region] Key as: Trionymus calamagrostidis
- Tang1992: pp.152 ( Adult (F) ) [Palaearctic region]
- Tang1992: pp.153 ( Adult (F) ) [Palaearctic region]
- Tang1992: pp.152 ( Adult (F) ) [Palaearctic region] Key as: Trionymus tomlini
- Danzig1988: pp.704 ( Adult (F) ) [Far East Russia]
- KosztaKo1988F: pp.159 ( Adult (F) ) [Central Europe]
- KosztaKo1988F: pp.159 ( Adult (F) ) [Central Europe] Key as: Trionymus tomlini
- Danzig1980a: pp.178 ( Adult (F) ) []
- Terezn1975: pp.263 ( Adult (F) ) [Ukraine]
- TerGri1973: pp.62 ( Adult (F) ) [Armenia]
- Willia1962: pp.59 ( Adult (F) ) [Great Britain]
- Willia1962: pp.58 ( Adult (F) ) [Palaearctic region] Key as: Trionymus tomlini
- Borchs1949: pp.95 ( Adult (F) ) [Palaearctic region] Key as: Trionymus calamagrostidis
- Borchs1949: pp.95 ( Adult (F) ) [Palaearctic region] Key as: Pseudococcus perrisii
Remarks
- Systematics: Type females of T. orientalis Matesova, 1968, were found to differ from females from Europe and Siberia. However, some individual variability was found within the type series: 7 specimens, including holotype, demonstrate the mentioned characters of T. orientalis, whereas one specimen resembled usual T. perrisii. Danzig and Gavrilov-Zimin determined that body pores and ducts vary widely in number in this species, sometimes in the insects from the same locality. Trionymus tomlini Green, 1925 was regarded as different from T. perrisii only in the presence of a larger circulus with an intersegmental fold. Danzig & Gavrilov-Zimin stated that the size and structure of circuli differ widely in this species and the character cannot be used for the species separation. Although the type material of P. pseudoperrisii was lost, this species did not differ from T. perrisii in the original description and figure by Rasina, 1971 and that they structure of cerarii and distribution of multilocular and tubular ducts lie within the range of variation in T. perrisii. They determined that since Pseudococcus calamagrostidis differed from T. perrisii only in having more numerous multilocular pores along the margin of the ventral sephalothorax and in the absence of a circulus, these characters could not be considered as significant to separate the species. Danzig & Gavrilov-Zimin (2015) compared syntypes of Trionymus dactylis Green, 1925 with T. perrisii and were unable to find any differences between the two. They also stated that they diagnostic character used in the key of Williams (1962), a smaller number of dorsal multilocular pores in T. dactylis, lies within the range of individual variation of T. perrisii.
- Structure: Female body elongate oval, up to 5 mm long, yellow, pink or violet in life. Antennae 8-segmented. Legs normally developed; hind coxae with numerous translucent pores. Anal apparatus complete. Both pairs of ostioles well developed. Circuli 1-5 in number; commonly only one small circulus present; if circuli 3-5 in number, they are significantly larger; sometimes circuli totally absent. (Danzig & Gavrilov-Zimin, 2015)
- Biology: Living at the base of the leaf sheaths of grasses. Adult females migrate from grasses to oviposit under the bark of dead oak branches in England. Life history and natural enemies discussed by Kosztarab & Kozár (1988F). Parasites in Palaearctic region discussed by Trjapitzin (1989).Living on roots of the host plant. Description and illustration of adult female of junior synonym, T. tomlini by Williams (1962), Tang (1992) and by Kwon et al. (2003a).
- General Remarks: Good description and illustration of the adult female given by Marchal (1908), Schmutterer (1952a), Williams (1962), Matesova (1968a) (as T. orientalis Matesova), Ter-Grigorian (1973), Tereznikova (1975) and by Danzig (1980b). Good description of the adult female given by Borchsenius (1949), Kosztarab & Kozár (1988F) and by Tang (1992). Koteja (1969) discussed the variation of several taxonomic characters. Good description and illustration of the adult female given by Danzig (1997a). Good description of the adult female given by Borchsenius (1949) and by Tang (1992).
Illustrations
Citations
- Balach1930: distribution, host, 313
- BatsanKaKi2017: distribution, host, 266
- BenDov1994: catalog, 523
- BenDov1994: catalog, 408-409, 508-509, 513-515, 523-524, 530-531
- Berg1899: taxonomy, 78
- BoratyWi1964a: distribution, host, 108
- Borchs1949: description, distribution, host, taxonomy, 142-146
- BugajNJuKa2021: distribution, host, list, 313
- CebeciAr2006: distribution, host, 144
- ChoiLe2022: phylogeny, 358
- Cocker1897: distribution, host, 201
- Danzig1959: distribution, host, 444
- Danzig1975: distribution, host, 48
- Danzig1980b: description, distribution, host, illustration, taxonomy, 178-179
- Danzig1997a: description, distribution, host, taxonomy, 96-99, 101-103
- DanzigGa2015: description, distribution, host, illustration, key, taxonomy, 164-171
- ESKSAE1994: distribution, list, 111
- Fernal1903b: taxonomy, 103
- Foldi2000: distribution, host, 80
- Foldi2001: distribution, 303-308
- FoldiGe2018: distribution, list, 13
- FoldiGe2018: distribution, list, 13
- FrancoRuMa2011: distribution, 21,25
- Gavril2003a: distribution, host, taxonomy, 108
- Gavril2016: distribution, host, 6
- GavrilKu2005: distribution, taxonomy, 19-34
- GavrilTr2008a: karyology, taxonomy, 74
- GermaiMaKa2015: taxonomy, 580
- Gertss2005a: distribution, host, 37
- GolanLaJa2001: host, taxonomy, 229-249
- Goux1931: distribution, host, 4
- Goux1933a: distribution, host, 236
- Goux1941a: description, distribution, host, taxonomy, 38
- Goux1941a: description, taxonomy, 37
- Green1915a: distribution, host, 178
- Green1920: description, distribution, host, taxonomy, 120, 121
- Green1926a: distribution, host, taxonomy, 183
- Hellri2004: distribution, host, 84
- Jansen2001: distribution, host, 197-206
- KaydanKiKo2005: distribution, host, 92
- KaydanKo2010a: distribution, host, 38
- KaydanKoYa2001: distribution, host, 381
- KaydanUlEr2007: catalog, distribution, host, 103
- Kiritc1936: distribution, host, 72
- Kiritc1940: description, distribution, host, taxonomy, 123
- Kohler2009a: distribution, host, 24
- KohlerEi2005: distribution, host, 161
- KohlerEi2006: distribution, host, 14
- KosztaKo1988F: description, distribution, host, illustration, taxonomy, 164-165, 169
- Koteja1969a: distribution, host, 5-7
- Koteja1974b: distribution, taxonomy, 75
- Koteja2000a: distribution, 172
- KotejaZa1969: distribution, host, 361-362
- KotejaZa1979: distribution, 671-672
- KotejaZa1983: distribution, host, 475
- Kozar1983a: distribution, host, 142
- Kozar1985: distribution, host, 201-205
- KozarDr1991: distribution, host, 362
- KozarKoSc1999: distribution, host, 111-114
- KozarOrKo1977: distribution, host, 71
- KozarSzFe2013: distribution, list of species, 93, 108-109, 126-127, 145, 163-164, 187, 199
- KozarTrPe1984: distribution, host, 5
- KozarTzVi1979: distribution, host, 130
- KozarWa1985: catalog, taxonomy, 73
- KozarWa1986: distribution, host, 115
- KozarzRe1975: distribution, host, 15-16
- Kraatz1888: taxonomy, 176
- Kraatz1888a: taxonomy, 360
- KwonDaPa2003a: distribution, host, taxonomy, 420-423
- Lagows1996: distribution, host, 31
- LagowsGo2020: distribution, 71
- LiuNiLu2022: phylogenetics,
- LongoMaPe1995: distribution, 121
- MalumpBa2012: distribution, 19
- Marcha1908: description, distribution, host, illustration, taxonomy, 248-251
- Marott1987a: distribution, host, 113
- Mateso1968: distribution, host, taxonomy, 103-105
- Mateso1968a: description, distribution, host, illustration, taxonomy, 152-154
- MazzeoLoRu1994: distribution, host, 205
- MazzeoRuLo1997: distribution, host, 21-22
- Newste1895: description, distribution, host, illustration, taxonomy, 167
- Ossian1985: distribution, host, 146
- Rasina1971: description, distribution, host, illustration, taxonomy, 3-6
- Reitte1898: taxonomy, 54
- Reyne1951: distribution, host, xxxix
- RussoMa1997: distribution, 45-55
- Schmut1952: description, distribution, host, illustration, taxonomy, 404-406
- Schmut1955b: biological control, distribution, host, 516,519
- SchmutHo2016: distribution, host, 32
- Seljak2010: distribution, host, 103
- Signor1875c: description, distribution, host, taxonomy, 337-338
- SimonKa2011: distribution, 236
- Stalaz2014: distribution, taxonomy,
- Stalaz2021: distribution, host, 453, 470, 471
- Strand1928: taxonomy, 47
- Suh2020: distribution, host, 8, 15
- SzitaFeBe2017: distribution, host, 473
- Tang1992: description, distribution, host, taxonomy, 156-157, 166-167, 177-178, 183, 766
- TangHaTa1992: distribution, host, 4
- Tao1999: distribution, host, taxonomy, 30
- TerGri1966: distribution, host, 84-85
- TerGri1969: distribution, host, 53
- TerGri1973: description, distribution, host, illustration, taxonomy, 66-67, 67-69
- Terezn1975: distribution, host, illustration, taxonomy, 265, 267-268
- TrenchTo2014: distribution, 67
- Trjapi1989: biological control, 120, 141, 219, 282,
- UlgentErYa2022: distribution, host, S126
- Willia1962: description, distribution, host, illustration, taxonomy, 63-67, 72-74
- Willia2017a: catalog, list of species, 246, 247
- Willia2017a: catalog, list of species, 250
- WilliaBe2015: taxonomy, 164, 166
- YerlikBaKa2021: key, 327