Valid Names Results

Heterococcus nudus (Green, 1926) (Pseudococcidae: Heterococcus)

Nomenclatural History

Phenacoccus nudus Green 1926: 172. Type data: ENGLAND: Camberley, from leaf sheaths of grasses. Syntypes, female, by subsequent designation Type depository: London: The Natural History Museum, England, UK; accepted valid name Illustr.
Heterococcus nudus (Green, 1926); Green 1928: 10. change of combination
Phenacoccus (Heterococcus) nudus Green, 1926; Goux 1933: 235. change of combination
Heterococcus tritici; Borchsenius 1937: 55. misidentification
Heterococcus graminicola Morrison 1945: 48. Type data: U.S.A.: Ohio, Wooster, on Phleum pratense. Holotype, female, Type depository: Washington: United States National Entomological Collection, U.S. National Museum of Natural History, District of Columbia, USA; junior synonym (discovered by Miller1975DR, 11).
Heterococcus occidentalis Morrison 1945: 53. Type data: U.S.A.: Washington, Yakima, on grass. Syntypes, female, Type depository: Washington: United States National Entomological Collection, U.S. National Museum of Natural History, District of Columbia, USA; junior synonym (discovered by Miller1975DR, 11).
Heterococcus borkhsenii Morrison 1945: 48. Type data: UKRAINE: Odessa, in leaf sheaths of wheat grass. Lectotype, female, by subsequent designation (Danzig1980b,143). Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; junior synonym (discovered by Danzig1968, 502). Illustr.
Heterococcus graminicolus Morrison, 1945; Ferris 1953a: 367. change of combination
Heterococcus variabilis Schmutterer 1958: 18. Type data: GERMANY: Oberammergau, on Agrostis alba. Holotype, female, Type depository: Wetlenberg: The Schmutterer Collection, Germany; junior synonym (discovered by Miller1975DR, 11).
Heterococcus pulverarius; Williams 1961: 673. misidentification (discovered by Miller1975DR, 11).
Heterococcus pulverarius; Williams 1962: 31. misidentification (discovered by Miller1975DR, 11).
Heterococcus pulverarius; Williams 1963: 101. misidentification (discovered by Miller1975DR, 11).
Heterococcus borchsenii Morrison, 1945; Ter-Grigorian 1966: 90. misspelling of species epithet
Heterococcus pulverarius; McKenzie 1967: 191. misidentification (discovered by Miller1975DR, 11).
Heterococcus pulverarius; Komosinska & Podsiadlo 1967: 684. misidentification (discovered by Miller1975DR, 11).
Heterococcus pulverarius; Danzig 1968: 502. misidentification (discovered by Miller1975DR, 11).
Heterococcus pulverarius; Koteja & Zak-Ogaza 1969: 360. misidentification (discovered by Miller1975DR, 11).
Heterococcus agropyri Savescu 1985: 119. Type data: ROMANIA: Bucarest, on Agropyron repens. Syntypes, female, by subsequent designation Type depository: Bucarest: Academie des Sciences Agricoles et Forestieres, Romania; junior synonym (discovered by DanzigGa2012a, 784). Illustr.

Common Names

naked grass-mealybug Koszta1996

Ecological Associates

Hosts:

Families: 3 | Genera: 31

Asteraceae

Haplopappus | BenDov1994 KozarHuFo1989

Pinaceae

Picea abies | BugajNJuKa2021

Poaceae

Agropyron | BenDov1994 KaydanKiKo2005 KozarHuFo1989
Agrostis | KozarGuBa1994
Agrostis capillaris | BugajNJuKa2021
Alopecurus | BenDov1994 Miller1975DR
Arrhenatherum elatius | BenDov1994 Miller1975DR
Avena sterilis | YerlikBaKa2023
Briza media | BenDov1994 Miller1975DR
Bromus | KozarGuBa1994
Bromus carinatus | BenDov1994 Miller1975DR
Bromus inermis | BenDov1994 Miller1975DR
Corynephorus canescens | BugajNJuKa2021
Cynodon dactylon | YerlikBaKa2023
Dactylis glomerata | Foldi2000
Danthonia decumbens | BenDov1994 Miller1975DR | (= Sieglingia decumbens)
Deschampsia cespitosa | BugajNJuKa2021
Echinochloa | KaydanKiKo2005
Echinochloa crus-galli | TongaAtBa2021
Elymus glaucus | BenDov1994 Miller1975DR
Elymus repens | BenDov1994 KaydanKiKo2005 KozarGuBa1994 KozarWa1986 Miller1975DR Savesc1985 | (= Agropyron repens)
Festuca airoides | BenDov1994 Danzig1980b | (= Festuca supina)
Festuca arundinacea | KaydanKiKo2005
Festuca cinerea | BugajNJuKa2021
Festuca ovina | BugajNJuKa2021
Festuca rubra | BenDov1994 KaydanKiKo2005 Miller1975DR
Holcus | BenDov1994 Miller1975DR
Holcus lanatus | BenDov1994 Foldi2000 Goux1933a
Hordeum | BenDov1994 TerGri1966
Hordeum murinum | BenDov1994 KaydanKiKo2005 Miller1975DR
Koeleria macrantha | BenDov1994 Danzig1980b | (= Koeleria gracilis)
Lolium | BenDov1994 KaydanKiKo2005 TerGri1966
Lolium perenne | Foldi2000 KaydanKiKo2005 KozarGuBa1994 Miller1975DR
Molinia caerulea | BenDov1994 Schmut1958
Panicum | BenDov1994 Miller1975DR
Phalaris canariensis | YerlikBaKa2023
Phleum pratense | BenDov1994 Miller1975DR Morris1945
Poa compressa | BugajNJuKa2021
Poa nemoralis | BenDov1994 Miller1975DR Schmut1958 | (= Agrostis alba)
Poa pratensis | BenDov1994 KotejaZa1983
Poaceae | Goux1931 Koszta1996 KozarTrPe1984 Marott1987a TerGri1966
Setaria viridis | BenDov1994 KaydanKiKo2005 Miller1975DR
Sorghum | KaydanKiKo2005 MyartsKaPo1995
Sorghum halepense | PellizPoSe2011
Stipa | KaydanKiKo2005
Zea mays | TongaAtBa2021

Foes:

Families: 1 | Genera: 2

Encyrtidae

Anusia tornike | JaposhYeKa2023
Cerchysius subplanus | JaposhYeKa2023

Associates:

Families: 1 | Genera: 1

Betaproteobacteria

Candidatus Tremblaya phenacola | ChoiLe2022

Geographic Distribution

Countries: 21

Armenia | BenDov1994 Borchs1949 TerGri1966

Azerbaijan | BenDov1994 Borchs1949

Canada

British Columbia | BenDov1994 KozarHuFo1989

Crete | PellizPoSe2011

France | BenDov1994 Foldi2000 Foldi2001 Goux1931 Goux1933a

Georgia (Republic of) | BenDov1994 Borchs1949

Germany | BenDov1994 Schmut1958

Greece | MilonaKo2008 MilonaKoKo2008a

Hungary | BenDov1994 KozarKiSa2004 KozarKo2002b KozarKoSc1999 KozarWa1986

Italy | BenDov1994 KosztaKo1988F KozarTrPe1984 LongoMaPe1995 Marott1987a

Moldova | BenDov1994 KozarOs1987

Netherlands | BenDov1994 Schmut1958

Poland | BenDov1994 GolanLaJa2001 Koteja1974b KotejaZa1983 Lagows1996 ZakOgaKo1964

Romania | BenDov1994 Savesc1985

Russia

Voronoezh Oblast | Gavril2003a
Yakutia-Sakha (=Yakut) AR | BenDov1994 Danzig1978a

Switzerland | KozarGuBa1994

Turkey | KaydanKiKo2005 KaydanUlEr2007

Turkmenistan | MyartsKaPo1995

Ukraine | BenDov1994 Borchs1949 Miller1975DR

Krym (=Crimea) Oblast | BenDov1994 Borchs1949

United Kingdom

England | BenDov1994 Green1926

United States

California | BenDov1994 McKenz1967 Miller1975DR
District of Columbia | BenDov1994 Miller1975DR
Idaho | BenDov1994 Miller1975DR
Kansas | BenDov1994 Miller1975DR
Maryland | Koszta1996 Miller1975DR
Massachusetts | Koszta1996
Michigan | Koszta1996
New Hampshire | Koszta1996 Miller1975DR
New Jersey | Koszta1996 Miller1975DR
New York | Koszta1996 Miller1975DR
North Dakota | BenDov1994 Miller1975DR
Ohio | Koszta1996 Miller1975DR
Oregon | BenDov1994 Miller1975DR
Pennsylvania | Koszta1996 Miller1975DR
Virginia | Koszta1996 Miller1975DR
Washington | BenDov1994 Miller1975DR
West Virginia | Koszta1996

Keys

DanzigGa2014: pp.610-612 ( Adult (F) ) [Palaearctic Heterococcus species]
DanzigGa2013: pp.460 ( ) [A Key to the Species of the Genus Heterococcus]
DanzigGa2012a: pp.784 ( Adult (F) ) [Heterococcus in the Palaearctic Region ]
Koszta1996: pp.135 ( Adult (F) ) [Northeastern North USA]
KosztaKo1988F: pp.105 ( Adult (F) ) [Central Europe]
Miller1975DR: pp.3 ( Adult (F) ) [World]
Miller1975DR: pp.35 ( Adult (M) ) [World]
Miller1975DR: pp.19, 24 ( Immature (F) ) [World]
Terezn1975: pp.190 ( Adult (F) ) [Ukraine & Palaearctic region]
McKenz1967: pp.189 ( Adult (F) ) [U.S.A., California]
Ferris1953a: pp.364 ( Adult (F) ) [North America]
Borchs1949: pp.267 ( Adult (F) ) [Palaearctic region]
Morris1945: pp.45 ( Adult (F) ) [World]

Remarks

Systematics: This species has frequently been misidentified as Trionymus pulverarius (Newstead) or Heterococcus pulverarius (now in Brevennia) (see Miller, 1975). Data on the taxonomy, synonyms, nomenclature, geographical distribution, host plants, biology and economic importance given by Ben-Dov (1994).
Structure: See colour illustration in McKenzie (1967).
Biology: Occurring in the leaf sheath and crown of its host plants.
Economic Importance: Caused browning and stunting of grasses in U.S.A., Washington state (Kosztarab, 1996).
General Remarks: Description and illustration of the adult female by Borchsenius (1949) (as H. borkhsenii), Ferris (1953a) (as H. graminicolus, Williams (1961) (as H. pulverarius), McKenzie (1967) (as H. pulverarius), Ter-Grigorian (1973), Miller (1975), Danzig (1980b) and by Kosztarab (1996). Good description and illustration of the adult female given by Savescu (1985).

Illustrations

Citations

BatsanKaKi2017: distribution, 263
BenDov1994: catalog, 191, 193-194
Borchs1937: description, distribution, host, illustration, taxonomy, 55
Borchs1949: description, distribution, host, taxonomy, 269
BugajNJuKa2021: distribution, host, list, 308
ChoiLe2022: phylogeny, 358, 359
Danzig1968: taxonomy, 502, 503
Danzig1978a: distribution, host, 74
Danzig1980b: description, distribution, host, illustration, taxonomy, 143-145
DanzigGa2012a: distribution, illustration, taxonomy, 784-786
DanzigGa2013: description, distribution, host, illustration, taxonomy, 460,462
DanzigGa2014: description, distribution, host, illustration, key, taxonomy, 620-622
DowellGiJe2016: distribution, 119
Ferris1953a: description, distribution, host, illustration, taxonomy, 367-368
Foldi2000: distribution, host, 78
Foldi2001: distribution, 303-308
FoldiGe2018: distribution, list, 12
Gavril2003a: distribution, host, 111
GavrilKu2005: distribution, taxonomy, 17-34
GavrilTr2008a: karyology, taxonomy, 74
GolanLaJa2001: distribution, host, taxonomy, 229-249
Goux1931: distribution, host, 3
Goux1933a: distribution, host, 235
Green1926a: description, distribution, host, illustration, taxonomy, 172, 173
Green1928: distribution, host, 10
HardyGuHo2008: molecular data, phylogeny, taxonomy, 51-71
HodgsoHa2013: phylogeny, taxonomy, 797
Jansen2001: distribution, host, 197-206
KaydanKiKo2005: distribution, host, 88
KaydanKo2010a: distribution, host, 34
KaydanUlEr2007: catalog, distribution, host, 100
KondoWa2022a: distribution, host, list, 23
Koszta1996: description, distribution, economic importance, host, illustration, taxonomy, 135, 136
KosztaKo1988F: description, distribution, host, illustration, taxonomy, 105-107
Koteja1974b: distribution, taxonomy, 74
Koteja2000a: distribution, 172
KotejaZa1983: distribution, host, 472
KozarGuBa1994: distribution, host, 153
KozarHuFo1989: distribution, host, 72
KozarKiSa2004: distribution, 57
KozarKo2002b: distribution, host, 375
KozarKoSc1999: distribution, host, 111-114
KozarOs1987: distribution, host, 92
KozarWa1986: distribution, host, 115
Lagows1996: distribution, host, 31
LagowsGo2020: distribution, 70
LongoMaPe1995: distribution, 118
Marott1987a: distribution, host, 109
McKenz1967: description, distribution, host, illustration, taxonomy, 191-193
Miller1975DR: description, distribution, host, illustration, taxonomy, 11-14
MilonaKo2008: distribution, host, 35-38
MilonaKoKo2008a: distribution, 143-147
Morris1945: description, distribution, host, illustration, taxonomy, 48-53
MyartsKaPo1995: distribution, host, 223-225
PellizChMi2015: distribution, 63,67
PellizPoSe2011: distribution, host, 292,296
RossHaOk2012: phylogeny, taxonomy, 199
Savesc1985: description, distribution, host, illustration, taxonomy, 119-121
Schmut1958:
SimonKa2011: distribution, 235
Teodor2018: distribution, history, 46
TerGri1966: distribution, host, 90
TerGri1973: description, distribution, host, illustration, taxonomy, 199-200
Terezn1975: description, distribution, host, illustration, taxonomy, 190, 191, 193
TongaAtBa2021: distribution, host, natural enemies,
UlgentErYa2022: distribution, host, S123
Willia1961: description, distribution, host, illustration, taxonomy, 673-675
Willia1963: description, distribution, host, taxonomy, 101
Willia2017a: catalog, list of species, 248
WilliaMa2012: distribution, taxonomy, 83
YerlikBaKa2023: distribution, host, illustration, list, 3, 4
ZakOgaKo1964: distribution, host, 423