Valid Names Results

Balanococcus kwoni is now Trionymus hamberdi (Borchsenius, 1949) (Pseudococcidae: Trionymus)

Nomenclatural History

Pseudococcus hamberdi Borchsenius 1949: 154. Type data: ARMENIA: Ashtarak District, Archashan near Amberd, on leaf sheaths of Phragmites sp.. Lectotype, female, by subsequent designation (Danzig1997a,103). Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; accepted valid name
Pseudococcus gracilipes Borchsenius 1949: 155. Type data: RUSSIA: North Caucasus, Maykop District, Kamennomostskaya, bank of Belaya River, on grass.. Lectotype, female, by subsequent designation (Danzig1997a,103). Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; junior synonym (discovered by Danzig1997a, 103).
Trionymus orestes Williams 1962: 63. Type data: ENGLAND: Surrey, Box Hill, on Brachypodium sylvaticun. Holotype, female, by original designation Type depository: London: The Natural History Museum, England, UK; junior synonym (discovered by DanzigGa2015, 133). Illustr.
Trionymus hamberdi (Borchsenius, 1949); Ter-Grigorian 1966: 85. change of combination
Trionymus gracilipes (Borchsenius, 1949); Kozár & Walter 1985: 73. change of combination
Trionymus subradicum Danzig 1985: 120. Type data: RUSSIA: Karachay-Cherkessia AR, Teberda, in leaf axils of timothy grass (Phleum pratense). Holotype, female, Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; junior synonym (discovered by Danzig1997a, 103). Illustr.
Balanococcus bambusum; Kwon, Danzig & Park 2003a: 398. misidentification (discovered by PellizDa2007, 65).
Balanococcus kwoni Pellizzari & Danzig 2007: 65. Type data: ITALY: Padua, Botanic Gardens, on Pseudosasa japonica; collected G. Pellizzari 16.xi.2006.. Holotype, female, by original designation Type depository: Padova: Dipartimento Agronomia Ambientale Produzioni Vegetali - Entomologia, Italy; junior synonym (discovered by DanzigGa2015, 133). Illustr.

Common Names

Hamberd's mealybug

Ecological Associates

Hosts:

Families: 2 | Genera: 25

Cyperaceae

Carex leporina | BenDov1994 Danzig1985

Poaceae

Agropyron | BenDov1994 Borchs1949 TerGri1966
Agrostis capillaris | BenDov1994 Danzig1985 | (= Agrostis vulgaris)
Anthoxanthum odoratum | BugajNJuKa2021
Bambusa | MalumpBa2012
Brachypodium sylvaticum | BenDov1994 Willia1962
Calamagrostis | BugajNJuKa2021
Calamagrostis arundinacea | BenDov1994 Danzig1985
Chimonocalamus | UlgentPoPe2014
Corynephorus | BugajNJuKa2021
Corynephorus canescens | BugajNJuKa2021
Deschampsia cespitosa | BenDov1994 Danzig1985
Elymus repens | BenDov1994 Danzig1985 | (= Agropyron repens)
Fargesia | LiWu2014
Fargesia nitida | LiTsWu2014 MalumpBa2012 | (= Sinarundinaria nitida)
Festuca ovina | BenDov1994 Danzig1985
Holcus lanatus | BenDov1994 Danzig1985
Hordeum | BenDov1994 TerGri1966 TerGri1973
Indocalamus longiauritus | LiWu2014
Indocalamus tessellatus | LiWu2014
Koeleria glauca | BugajNJuKa2021
Pharus | MalumpBa2012
Phragmites | BenDov1994 Borchs1949 TerGri1966 TerGri1973
Phyllostachys | MalumpBa2012
Phyllostachys bambusoides | MalumpBa2012
Phyllostachys nigra | LiTsWu2014 MalumpBa2012
Phyllostachys praecox | LiTsWu2014
Phyllostachys violascens | LiWu2014
Phyllostachys vivax | LiTsWu2014 MalumpBa2012 | (= Bambusa vivax)
Pleioblastus fortunei | MalumpBa2012
Pleioblastus linearis | MalumpBa2012
Pleioblastus variegatus | LiTsWu2014
Poa palustris | BenDov1994 Danzig1985
Poaceae | BenDov1994 Borchs1949
Pseudosasa japonica | PellizDa2007
Thamnocalamus spathiflorus | MalumpBa2012 | (= Thamnocalamus crassinodus.)

Geographic Distribution

Countries: 10

Armenia | Borchs1949 TerGri1966

China

Hubei (=Hupei) | LiWu2014
Zhejiang (=Chekiang) | LiWu2014

France | MatileWi2015

Hungary | Kozar1980 KozarKiSa2004 KozarKo2002b

Italy | Pelliz2010 PellizDa2007 | LongoMaPe1995

Kazakhstan | Danzig1997a

Poland | Danzig1985 GolanLaJa2001 Lagows1996

Russia

Caucasus | Borchs1949
Irkutsk Oblast | Danzig1997a
Kaliningrad Oblast | Danzig1997a
Karachay-Cherkessia AR | Danzig1985
St. Petersburg (=Leningrad) Oblast | Danzig1985 Danzig1997a

South Korea | UlgentPoPe2014

United Kingdom

England | MalumpBa2012
England | BenDov1994 Willia1962

Keys

YerlikBaKa2021: pp.326-327 ( Adult (F) ) [Trionymus of the Palearctic region]
DanzigGa2015: pp.99-106 ( Adult (F) ) [Palaearctic Trionymus species]
LiWu2014: pp.270 ( Adult (F) ) [Key to Balanococcus species of China] Key as: Balanococcus kwoni
LiTsWu2014: pp.371 ( Adult (F) ) [Mealybugs with legs on bamboo in China] Key as: Balanococcus kwoni
Danzig1997a: pp.96 ( Adult (F) ) [Palaearctic region]
Tang1992: pp.152 ( Adult (F) ) [Palaearctic region]
Danzig1988: pp.704 ( Adult (F) ) [Far East Russia]
KosztaKo1988F: pp.159 ( Adult (F) ) [Central Europe]
TerGri1973: pp.63 ( Adult (F) ) [Armenia]
Willia1962: pp.59 ( Adult (F) ) [British Pseudococcidae] Key as: Trionymus orestes
Borchs1949: pp.96 ( Adult (F) ) [Palaearctic region] Key as: Pseudococcus hamberdi

Remarks

Systematics: This species has been mis-identified as Balanococcus banbusum by Kwon, Danzig & Park (2003a). Danzig & Gavrilov-Zimin (2015) reviewed the mixed type material of Balanocccus kwoni and determined that the females from Italy (holotype ad some paratypes) did not differ from the types of Trionymus hamberdi, whereas females from Korea were conspecific with Trionymus orientalis.
Structure: Female body elongate, up to 4 mm long, pink in life. Antennae 7-8 segmented. Legs thin; hind coxae with translucent pores. Anal apparatus complete. Both pairs of ostioles present. Circulus small, from oval to almost quadrate. (Danzig & Gavrilov-Zimin, 2015)
Biology: Lives in leaf sheaths of grasses. Junior synonym, Balanococcus kwoni described as living exclusively on bamboo. (Williams and Malumphy, 2012)
General Remarks: Good description and illustration of the adult female given by Ter-Grigorian (1973), Danzig (1985) (as T. subradicum Danzig) and by Danzig (1997a). Good description of the adult female given by Borchsenius (1949) and by Tang (1992). Good description and illustration of the adult female of junior synonym, Trionymus orestes, given by Williams (1962). Description and illustration of adult female of junior synonym, Balanococcus kwoni, by Pellizzari & Danzig (2007).

Illustrations

Citations

AhnKiKi2020: distribution, host, 8
BenDov1994: catalog, 513-514, 522
Borchs1949: description, distribution, host, taxonomy, 154-155
BugajNJuKa2021: distribution, host, list, 312
Danzig1985: description, distribution, host, illustration, taxonomy, 119-121
Danzig1997a: description, distribution, host, illustration, taxonomy, 103-104
Gavril2003a: distribution, host, 108
Gavril2018: reproduction, 227
GavrilKu2005: distribution, 19-34
GolanLaJa2001: distribution, host, taxonomy, 229-249
KalandSi2014: distribution,
KosztaKo1988F: description, distribution, host, taxonomy, 161-162
KozarKiSa2004: distribution, 58
KozarKo2002b: distribution, host, 375
KozarWa1985: catalog, taxonomy, 73
KwonDaPa2003a: taxonomy, 398
LagowsGo2020: distribution, 71
LagowsGo2020: distribution, 71
LagowsKo1996: distribution, host, 31, 34
LiTsWu2014: biology, distribution, host, key, taxonomy, 371,372
LiWu2014: description, distribution, host, illustration, structure, taxonomy, 270, 272-273
LongoMaPe1995: distribution, 121
Malump2011: host, 169
MalumpBa2012: distribution, economic importance, host, 18,38
MatileWi2015: distribution, host, 74
MazzeoLoPe2014: distribution, 58
Pelliz2010: distribution, host, 90
PellizDa2007: description, distribution, host, illustration, taxonomy, 65-68
SimonKa2011: distribution, 236
Suh2020: distribution, host, 17
Suh2020: distribution, host, 17
Tang1992: description, distribution, host, taxonomy, 166
TerGri1966: distribution, host, 85
TerGri1973: description, distribution, host, illustration, taxonomy, 70-72
UlgentPe2013: host, 493
UlgentPoPe2014: distribution, host, illustration, 77, 79, 80
Willia1962: description, distribution, host, illustration, taxonomy, 63-64
WilliaMa2012: distribution, 83,86
WilliaMa2012: distribution, 85
YerlikBaKa2021: key, 327