Valid Names Results
Dynaspidiotus californicus (Coleman, 1903) (Diaspididae: Dynaspidiotus)Nomenclatural History
- Aspidiotus (?) pini Comstock 1881a: 306. Type data: U.S.A.: New York, Ithaca, on leaves of Pinus rigida and Georgia, Macon, on Pinus mitis.. Syntypes, female, Type depository: Washington: United States National Entomological Collection, U.S. National Museum of Natural History, District of Columbia, USA; junior homonym Notes: Homonym of Aspidiotus pini Hartig (1839) and Bouche (1851). Illustr.
- Aspidiotus abietis; Cockerell 1894l: 190. misidentification (discovered by Borchs1966, 275).
- Aspidiotus abietis pini Comstock, 1881; Cockerell 1896b: 333. change in status (level)
- Aspidiotus californicus Coleman 1903: 64. Type data: U.S.A.: California, at nine localities, on Pinus sabiniana, P. ponderosa, P. lambertiana and P. attenuata.. Syntypes, female, Type depository: Davis: The Bohart Museum of Entomology, University of California, California, USA; junior synonym replacing a junior homonym Notes: Replacement name for Aspidiotus ? pini Comstock, 1881a.
- Nuculaspis californica (Coleman, 1903); Ferris 1938a: 251. change of combination requiring emendation of specific epithet for agreement in gender
- Aspidiotus (Nuculaspis) californicus Coleman, 1903; Merrill 1953: 16. change of combination
- Dynaspidiotus californicus (Coleman, 1903); Danzig 1993: 149. change of combination
- Nuculaspis californica (Coleman, 1903); Miller & Davidson 2005: 296. revived combination (previously published)
- Dynaspidiotus californica (Coleman, 1903); Normark 2019: S5. misspelling of species epithet
Common Names
- black pine leaf scale Dekle1965c McKenz1956 Merril1953
- escama negra de las hojas de pino KondoWa2022b
Ecological Associates
Hosts:
Families: 1 | Genera: 2
- Pinaceae
- Pinus | BesheaTiHo1973 Dekle1965c Ferris1920b
- Pinus attenuata | Colema1903 Sander1906
- Pinus cembroides | McDani1970 McKenz1956
- Pinus contorta | Ferris1938a McKenz1956 | (= Pinus murrayana)
- Pinus coulteri | NormarOkMo2019
- Pinus echinata | Comsto1881a McKenz1956 | (= Pinus mitis)
- Pinus lambertiana | Colema1903 McKenz1956 Sander1906
- Pinus monophylla | NormarOkMo2019
- Pinus occidentalis | GomezM1941
- Pinus palustris | MerrilCh1923
- Pinus ponderosa | Colema1903 McDani1970 McKenz1956 Sander1906
- Pinus radiata | Ferris1938a McKenz1956
- Pinus rigida | Comsto1881a McKenz1956
- Pinus sabiniana | Colema1903 McDani1970 McKenz1956 Sander1906
- Pinus tropicalis | MestreHaEv2011
- Pseudotsuga menziesii | Ferris1920b Ferris1938a McDani1970 McKenz1956 MerrilCh1923 | (= Pseudotsuga taxifolia)
Foes:
Families: 2 | Genera: 3
Geographic Distribution
Countries: 6
- Canada | Nakaha1982
- Cuba | MestreHaEv2011
- Dominican Republic | GomezM1941
- Haiti | PerezG2008
- Mexico | Ferris1938a
- United States
- Alabama | WaltmaRaWi2016
- California | Ferris1938a McKenz1956 Sander1906
- Florida | Dekle1965c Merril1953 MerrilCh1923
- Georgia | BesheaTiHo1973 Comsto1881a
- Nevada | NormarOkMo2019
- New York | Comsto1881a
- Oregon | SchuhMo1948
- Texas | Ferris1938a McDani1970
Keys
- MillerDa2005: pp.29 ( Adult (F) ) [Field Key to Economic Armored Scales] Key as: Nuculaspis californica
- Koszta1996: pp.543 ( Adult (F) ) [Northeastern North America]
- McKenz1956: pp.26 ( Adult (F) ) [U.S.A.: California]
- Ferris1942: pp.39 ( Adult (F) ) [North America]
- Britto1923: pp.371 ( Adult (F) ) [U.S.A.: Connecticut]
- Comsto1883: pp.55-57 ( Adult (F) ) [North America]
Remarks
- Systematics: Genbank accession numbers: KF887380 and KF887381. Aspidiotus ? pini Comstock, 1881a: 306, (described from Pinus in USA, New York) is a homonym of Aspidiotus pini Hartig, 1839 and Aspidiotus pini Bouche, 1851. No Replacement Name has been formally proposed for Aspidiotus ? pini Comstock, 1881. However, Aspidiotus californicus Coleman, 1903 (described from Pinus spp. in USA, California) was found by Ferris (1938a: 190) to be a synonym of Aspidiotus ? pini Comstock, 1881a, becoming therefore a Replacement Name.
- Structure: Female scale oblong-oval, and rather conical, 2 mm long, 1 mm wide; colour blackish with pale edges; exuviae central, reddish-brown; great variation in size and form were observed (Coleman, 1903). Scale of the female quite black, usually with one margin set against the edge of a needle and therefore the scale forced to be slightly elongate, exuviae central; scale of the male darker, elongate oval, exuvia central (Ferris, 1938a). Colour photograph by Gill (1997).
- Biology: Occurring on the needles (Ferris, 1938a).
- Economic Importance: The black pineleaf scale is considered a serious pest of pines in the West Coast of the USA (Gill, 1997).
- General Remarks: Description and illustration of adult female by Ferris (1920b; 1938a), McKenzie (1956), Kosztarab (1996) and by Gill (1997).
Illustrations
Citations
- Alstad1998: distribution, ecology, host, life history, 3-21
- AlstadEd1983: distribution, ecology, host, life history, 93-95
- AlstadEd1983a: distribution, ecology, host, life history, 413-426
- AlstadEd1987: distribution, ecology, host, life history, 652-654
- AlstadEd1989: distribution, ecology, host, life history, 253-263
- AlstadEdJo1980: distribution, ecology, host, life history, 665-667
- Amos1933: taxonomy, 208
- AndersWuGr2010: molecular data, 992-1003
- Baker1972: control, distribution, economic importance, host, 1-642
- BeardsDaHo1976: economic importance, 103
- BenDovGe2003: catalog, 478-480
- BesheaTiHo1973: distribution, host, 7
- Borchs1966: catalog, 275
- Britto1923: description, distribution, host, taxonomy, 371,372
- BrownEa1967: control, distribution, economic importance, host, 1-72
- CampbeLaHu2014: DNA, genebank, phylogeny, 531, 532, 533
- Cocker1894l: description, distribution, host, taxonomy, 190-191
- Cocker1896b: distribution, taxonomy, 333
- Colema1903: description, distribution, host, taxonomy, 64
- Comper1928: biological control, 209-230
- Comsto1881a: description, distribution, host, illustration, taxonomy, 306-307
- Comsto1883: taxonomy, 67
- DeSant1940: biological control, 29-44
- Dekle1965c: description, distribution, host, taxonomy, 98
- Dekle1976: description, distribution, economic importance, host, taxonomy, 119
- Drea1990: biological control, 51-59
- Edmund1973: biological control, distribution, ecology, host, life history, 765-777
- EdmundAl1958: distribution, ecology, host, life history, 391-392
- EdmundAl1981: distribution, ecology, host, life history,
- EdmundAl1984: distribution, ecology, host, life history, 267-268
- EdmundAl1985: chemical control, distribution, ecology, host, life history, 403-405
- Essig1928: distribution, host, 76-78
- Ferris1920b: description, distribution, host, illustration, taxonomy, 52-54
- Ferris1938a: description, distribution, host, illustration, taxonomy, 251
- Ferris1938b: taxonomy, 65,70
- Ferris1941e: taxonomy, 41,47
- Ferris1942: taxonomy, 446:39
- Frank2021: ecology, life history, 5
- FurnisCa1977: distribution, host, 111
- Gill1997: description, distribution, economic importance, host, illustration, taxonomy, 204-205,207
- GomezM1941: distribution, host, 128
- Gordh1979: biological control, 907
- Gould1983: ecology, life history, structure, 599
- GruwelMoNo2007: endosymbionts, taxonomy, 267-280
- HagenVaDa1971: biological control, distribution, host, 253-293
- Herric1911: description, distribution, host, illustration, taxonomy, 13
- Howard1895e: biological control, 1-44
- JohnsoYoAl1997: chemistry, distribution, ecology, host, life history, 1794-1804
- Keen1952: distribution, economic importance, host, 1-280
- KondoWa2022a: distribution, host, list, 16
- KondoWa2022h: control, diagnosis, distribution, host, illustration, taxonomy, 473-474
- Koszta1996: biological control, description, distribution, economic importance, host, illustration, life history, taxonomy, 543-544
- Kuwana1932: taxonomy, 54
- Leonar1897: taxonomy, 285
- Lindin1957: taxonomy, 550
- Lobdel1937: taxonomy, 78
- MacGil1921: description, distribution, host, taxonomy, 396
- McClur1990a: distribution, ecology, host, 165-168
- McClur1990g: distribution, ecology, host, taxonomy, 319-330
- McDani1970: distribution, host, illustration, taxonomy, 425-427
- McKenz1939: taxonomy, 53
- McKenz1956: description, distribution, host, illustration, taxonomy, 77-80
- Merril1953: description, distribution, host, taxonomy, 16-17
- MerrilCh1923: description, distribution, economic importance, host, taxonomy, 207-208
- MestreHaEv2011: distribution, host, 11-12
- Miller1983: distribution, host, 4-6
- MillerDa1990: distribution, economic importance, host, 304
- MillerDa2005: description, distribution, economic importance, host, illustration, taxonomy, 296-299
- MorseGrCl2005: molecular data, phylogeny, taxonomy, 79-94
- MorseNo2006: molecular biology, phylogeny, 338-349
- Nakaha1982: distribution, host, 61
- NormarOkMo2019: distribution, host, phylogeny, 20, S5, S6
- Nur1990a: chromosomes, structure, taxonomy, 185
- PerezG2008: distribution, 214
- Raushe1983: ecology, host, life history, 223-257
- RugmanAnMo2010: molecular data, phylogenetics, taxonomy, 30-38
- Sander1906: catalog, distribution, host, taxonomy, 13
- SchmutKlLu1957: distribution, economic importance, host, 494
- SchneiFiNo2019: key, taxonomy, 92
- SchneiOkNo2018: phylogeny,
- SchneiSkOc2019: economic importance, 324
- SchuhMo1948: diagnosis, distribution, host, 42
- StrublJo1964: distribution, economic importance, host, 1-6
- VonEll2025: distribution, 24
- WaltmaRaWi2016: distribution, 231
- Watson2001: list, 177
- Willia2017a: catalog, list of species, 219
- Wilson1917: description, distribution, host, taxonomy, 31-32
- Woodwo1903: taxonomy, 38
- YoungJoAl1993: distribution, ecology, host, life history, 497
- Zahrad1990: description, distribution, host, 641-642


