Takagi, S. 2002 One new subfamily and two new tribes of the Diasididae (Homoptera: Coccoidea).. Insecta Matsumurana (New Series) 59: 55-100.

Notes: Ulucoccinae, subfam. n. and Thysanaspidini and Smilacicolini, tribe, n., are erected on the basis of Ulucoccus, Thysanaspis, and Smilacicola, respectively. The subfamily Ulucoccinae is one of very primitive forms of the Diaspididae, lacking distinct pygidial appendages, and may approach a remote ancestor of the Diaspidinae, which is similar to the second instar males of some species of the tribe, especially of the Chionaspidina. The tribe Thysanaspidini represents the most primitive form of the Aspidiotinae in the pygidial appendages, which are composed of well-developed pectinae alone in the nymphal stages. Thysanaspis is pupillarial, so that the nymphal stages of the genus give some idea of the putative ancestor of the extant Aspidiotinae. The Smilacicolini inhabit a constricted space on the plant body as the Odonaspidini do, and may have derived from a group of forms related to, but more primitive than, the extant Parlatoriini, to which the Odonaspidini are also related. Five other diaspidids are decribed or figured for comparison with the Thysanaspidini and the Smilacicolini. Two of them, occurring in the Philippines, are new species belonging to a new genus: Ligaspis maculata and L. pala. Another species, Mongrovaspis quadrispinosa, is newly recorded from eastern Asia and is redescribed. The new higher taxa and the other higher taxa mentioned in this study are primarily gradal units, and comments are made on the reason why such a view of taxa is adopted.