Valid Names Results

Cornoculus is now Ovaticoccus Kloet, 1944 (Eriococcidae)

Nomenclatural History

• Gymnococcus Cockerell 1894v: 1053. Type species: Coccus agavium Douglas by monotypy . junior homonym (discovered by Kloet1944: 86) Notes: Douglas (1888c) stated, in his description of Coccus agavium: "At one time I thought it might constitute the type of a new genus, under the name of Gymnococcus, but in consideration of the important and leading characters of the antennae, I have concluded (for the present, at least) that it is better to regard all the others as specific, and to refer the species to Signoret's genus Coccus." Cockerell (1894v) designated Gymnococcus as a distinct genus and gave Douglas credit. Since Douglas did not actually use the name Gymnococcus, and since Cockerell was the first person to do so, Cockerell is here given credit for describing this genus (see also Hoy, 1963). Parrott (1900) considered Gymnococcus valid but credited it to Newstead (1897). However, Gymnococcus Douglas was considered valid until Kloet (1944) discovered that the name Gymnococcus was preoccupied by a genus of Mycetozoa (Gymnococcus Zopf, 1887) and introduced the new name Ovaticoccus. This name was disregarded until Ferris (1957c) confirmed Kloet's designation.
• Ovaticoccus Kloet 1944: 86. Type species: Coccus agavium Douglas by monotypy . replacement name
• Gymnococcus (Parrotia) Gómez-Menor Ortega 1954: 142. . unavailable name that is placed (discovered by MorrisMo1966: 149) Notes: According to Morrison & Morrison (1966) Gómez-Menor Ortega proposed Parrotia as a subgenus of Gymnococcus for two described American species whose name he did not include. Since no type species was given, it is unavailable according to Article 13 (b) of the 1961 code. Further, it is preoccupied by Parrotia Kieffer (1924) in the Diptera.
• Cornoculus Ferris 1955a: 81. Type species: Cornoculus cornutus Ferris by monotypy . junior synonym (discovered by MillerSt2022: 5)
• Onceropyga Ferris 1955a: 208. Type species: Eriococcus neglectus Cockerell by original designation . synonym and homonym (discovered by MillerSt2022: 5) Notes: Homonym of Onceropyga Turner, 1906 (Lepidoptera: Zygaenidae); discovered by Hoy, 1963: 179.
• Ovatococcus Lindinger 1957: 549;. . misspelling of genus name
• Ovaticoccus (Parrottia) Boratynski 1958;. . misspelling of genus name (discovered by MorrisMo1966: 149) Notes: Misspelling of the Subgenus name.
• Oregmopyga Hoy 1963: 179. . replacement name that is a synonym Notes: (replacement name for the junior homonym Onceropyga Ferris 1955)
• Ovatococcus Danzig 1971d: 821;. . misspelling of genus name

Remarks

• Systematics: Ovaticoccus is similar to Acanthococcus in having: enlarged setae; microtubular ducts each with two internal scleroses; claw digitules longer than claw and capitate. Species of Acanthococcus differ by (character states of Ovaticoccus species are given in brackets): anal lobes protruding, each lobe usually with three or four enlarged setae (anal lobes not protruding or slightly protruding, each lobe with two or fewer enlarged setae); anal ring with four setae on each side of ring and with at least one ring of cells (anal ring with zero to three setae on each side of ring and often with reduced number of cells); enlarged setae often conical, not pear- or nipple-shaped (enlarged setae sometimes conical, often pear- or nipple-shaped). Species of Ovaticoccus are similar also to species of the Old-World genus Hispaniococcus Kozár in having: three setae on each side of anal ring; enlarged setae nipple-shaped in one species of Hispaniococcus; and microtubular ducts with two internal sclerotized areas. Species ofHispaniococcus differ from species of Ovaticoccus by (character states in brackets are of Ovaticoccus): claw digitules shorter than length of claw and apically acute (claw digitules same length as, or longer than, length of claw and apically capitate); anal lobes protruding; and each lobe with three or more enlarged setae (anal lobes not protruding or slightly protruding, and each lobe with two or fewer enlarged setae). (Miller & Stocks, 2022) Generic characteristics that distinguish this genus from all other Eriococcids are: dorsal setae, when present, nipple shaped; without anal lobes (Miller & McKenzie, 1967). In Kozár, et al., 2013 Ovaticoccus was placed in the family Acanthococcidae Signoret, 1875, but is here kept in Eriococcidae because placement of species.outside of the palaearctic are unclear.

  Ferris (1955) characterized the single species of Cornoculus (C. cornutus) as having: unusually large hornlike eyes; characteristically shaped setae, e.g., nipple-shaped; antennae 7-segmented; multilocular pores on dorsum and venter; and anal ring reduced. Miller and McKenzie (1967) added a second species to Cornoculus (C. densus Miller), based primarily on the large number of macrotubular ducts on the lateral areas of the venter but not having unusual eyes. With the exception of the unusual eyes, the species of Cornoculus possess all of the characteristics of species of Ovaticoccus and are considered by Miller & Stocks (2022) to be part of the latter genus. The unusual large eyes are believed to be an autapomorphy.

• Structure: Adult females oval, body various shades of red, pink and purple. Species are normally host-specific and found under bark, but in sheaths when on Gramineae or Agavaceae. Exposed only when infestations are heavy (Miller & McKenzie, 1967).
• General Remarks: Description by González (2009) of the adult female.

Keys

• MillerSt2022: pp.11-12 ( Adult (F) ) [Nearctic genera of Eriococcidae]
• Hodgso2020: pp.19-22 ( Adult (M) ) [Neococcoid higher taxa]
• KondoGuCo2016: pp.475-476 ( Adult (F) ) [Genera of the Eriococcidae of South America] Key as: Oregmopyga
• KozarKaKo2013: pp.62-63 ( Adult (F) ) [Key to genera of Acanthococcidae]
• KozarKaKo2013: pp.62-63 ( Adult (F) ) [Key to genera of Acanthococcidae]
• PellizKo2011: pp.58 ( Adult (F) ) [Key to the genera of Eriococcidae in Europe]
• HodgsoMaMi2011: pp.54-55 ( Adult (F) ) [Key to the Eriococcidae of the Neotropical Region based on the morphology of adult females]
• HodgsoMaMi2011: pp.54-55 ( Adult (F) ) [Key to the Eriococcidae of the Neotropical Region based on the morphology of adult females] Key as: Oregmopyga
• HodgsoMaMi2011: pp.54-55 ( Adult (F) ) [Key to the Eriococcidae of the Neotropical Region based on the morphology of adult females]
• HodgsoMi2010: pp.90-92 ( First instar ) [Key to the Genera of the Eriococcidae of South America] Key as: Oregmopyga
• Miller2005: pp.491 ( Adult (F) ) [Genera of Eriococcidae of the Eastern U.S.] Key as: Oregmopyga
• Miller2005: pp.491 ( Adult (F) ) [] Key as: Genera of Eriococcidae of the Eastern U.S.
• KondoGuMi2004: pp.3 ( Adult (F) ) [Key to species] Key as: Oregmopyga
• TangHa1995: pp.642 ( Adult (F) ) [Key to genera of Calycicoccina] Key as: Gymnococcus, Ovaticoccus
• Gill1993: pp.171-172 ( Adult (F) ) [Oregmopyga species of CaliforniaKey to the California Genera of Eriococcidae] Key as: Oregmopyga
• MillerMi1993: pp.77 ( Adult (F) ) [Oregmopyga species of the eastern United States] Key as: Oregmopyga
• Gill1993: pp.173 ( Adult (F) ) [Ovaticoccus species of California, Key to California Genera of Eriococcidae]
• MillerMi1993: pp.6 ( Adult (F) ) [Key to Genera of Eriococcidae of the Eastern U.S.]
• MillerMi1993: pp.6 ( Adult (F) ) [Key to Genera of Eriococcidae of the Eastern U.S.] Key as: Oregmopyga
• Willia1985h: pp.352 ( Adult (F) ) [Key to genera of British Eriococcidae]
• Danzig1971d: pp.821 ( Adult (F) ) [Key to genera of Eriococcidae]
• MillerMc1967: pp.490 ( Adult (F) ) [North American species of Orgemopyga] Key as: Oregmopyga
• MillerMc1967: pp.507 ( Adult (F) ) [North American species of Ovaticoccus]
• Ferris1955a: pp.208 ( Adult (F) ) [North American species of Onceropyga] Key as: Onceropyga
• Ferris1955a: pp.179 ( Adult (F) ) [North American species of Gymnococcus] Key as: Gymnococcus
• Ferris1955a: pp.179 ( Adult (F) ) [North American species of Gymnococcus] Key as: Gymnococcus

Associated References

• Arnett1985: distribution, taxonomy, pp. 239
• Balach1942: taxonomy, pp. 42
• Balach1948b: taxonomy, pp. 254
• Boraty1958: description, distribution, taxonomy, pp. 173-175
• BoratyWi1964: taxonomy, pp. 92
• Borchs1949: taxonomy, pp. 322, 369
• Brown1967: distribution, host, pp. 132
• Cocker1894v: distribution, taxonomy, pp. 1053
• Cocker1896b: taxonomy, pp. 323
• Cocker1899m: taxonomy, pp. 277
• Cocker1902j: taxonomy, pp. 717
• Cocker1905b: taxonomy, pp. 192-193
• Danzig1964: distribution, taxonomy, pp. 632, 634
• Danzig1971d: taxonomy, pp. 821, 824
• Dougla1888c: taxonomy, pp. 150
• Ferris1919a: taxonomy, pp. 19
• Ferris1921b: taxonomy, pp. 60
• Ferris1922b: taxonomy, pp. 247
• Ferris1955a: description, taxonomy, pp. 178, 208
• Ferris1957b: taxonomy, pp. 66
• Ferris1957c: distribution, taxonomy, pp. 88
• Gill1993: distribution, taxonomy, pp. 171, 173
• GomezM1954: description, taxonomy, pp. 141
• Hodgso2020: key, pp. 20
• HodgsoMaMi2011: taxonomy, pp. 54-55
• HodgsoMi2010: description, illustration, taxonomy, pp. 66-70
• Hoy1962: taxonomy, pp. 13, 201
• Hoy1963: catalog, taxonomy, pp. 179, 181-182
• Kloet1944: taxonomy, pp. 86
• Kohler1998: catalog, distribution, taxonomy, pp. 393-394
• KondoGuCo2016: key, pp. 476
• KondoGuMi2004: taxonomy, pp. 1-11
• Koteja1974: taxonomy, pp. 297
• Koteja1974b: taxonomy, pp. 77
• KotejaZa1981: taxonomy, pp. 501, 502
• Kozar2009: distribution, host, taxonomy, pp. 111,114
• KozarKaKo2013: description, distribution, host, illustration, structure, taxonomy, pp. 373-382
• KozarKo2008a: taxonomy, pp. 148
• KozarWa1985: catalog, distribution, taxonomy, pp. 393
• Lawson1917: taxonomy, pp. 186
• Lindin1937: taxonomy, pp. 186
• Lindin1957: taxonomy, pp. 549
• MacGil1921: taxonomy, pp. 129, 141
• McKenz1964a: description, distribution, taxonomy, pp. 21, 25
• Miller1970: taxonomy, pp. 157
• Miller2005: distribution, taxonomy, pp. 491
• MillerGi2000: catalog, taxonomy, pp. 417-418, 426-427
• MillerMc1967: description, taxonomy, pp. 489-490, 506
• MillerMi1993: description, taxonomy, pp. 77, 84
• MillerSt2017: biology, diagnosis, distribution, key, pp. 807-808
• MillerSt2022: description, diagnosis, key, taxonomy, pp. 11, 85-87
• MorrisMo1966: taxonomy, pp. 138, 141
• Newste1897: taxonomy, pp. 12-13
• Newste1903: taxonomy, pp. 203-204
• Parrot1900: description, taxonomy, pp. 141
• PellizKo2011: description, distribution, host, structure, taxonomy, pp. 61
• PooleGe1997: distribution, pp. 355
• PopenoPa1900: taxonomy, pp. 135
• TangHa1995: taxonomy, pp. 642
• Terezn1981: taxonomy, pp. 13, 57
• Tranfa1976: taxonomy, pp. 129-130
• Willia1985h: taxonomy, pp. 384-385
• Zopf1887: taxonomy, pp.

32 Species