Family Pennygullaniidae

Pennygullania Koteja & Azar


Pennygullania Koteja & Azar, 2008: 158. Type species: Pennygullania electrina Koteja & Azar, by original designation.

GENERAL REMARKS: Description and definition, based on alate adult male, by Koteja & Azar (2008).

SYSTEMATICS: Pennygullania is the type genus of the family Pennygullaniidae Koteja & Azar (2008). Recently, Gavrilov-Zimin & Danzig (2012) synonymised Pennygullaniidae with Pseudococcidae, arguing that the characters used by Koteja & Azar (2008) to define Pennygullaniidae were insufficient to distinguish it from Pseudococcidae. Furthermore, the adult male of Pennygullania has two pairs of glandular pouches on the abdomen, as is the case for adult males in the pseudococcid subfamily Phenacoccinae. However, careful study of Koteja & Azarís (2008) illustration suggests that there is a ring of at least three pairs of simple eyes on heavily sclerotised ocular sclerites that almost meet on the top of the head, a condition very similar to that in Putoidae. Thus, Hodgson & Hardy, 2013, concluded that there was sufficient evidence to keep these families separate. Diagnosis of the type genus: Slim neococcid with two pairs of simple eyes, slender appendages bearing very long setae, solid scutum, trochanter and femur fused, two pairs of strong tail setae (with two setae in each group), penial sheath of coccid s. str. type.

KEYS: Koteja & Azar 2008: 135 (male) [Lower Cretaceous Lebanese amber inclusions].

CITATIONS: GavrilDa2012 [taxonomy: 108]; KotejaAz2008 [taxonomy, description: 158-161].

Pennygullania electrina Koteja & Azar


Pennygullania electrina Koteja & Azar, 2008: 159. Type data: LEBANON: Lower Cretaceous, Mdeyrij/Hammana, Caza Baabda, Mouhafazet Jabal Loubnan (Central Lebanon). Holotype male. Type depository: Paris: Museum National d'Histoire naturelle, France; type no. HAM-381. Described: male. Illust.

DISTRIBUTION: Palaearctic: Lebanon [KotejaAz2008].

GENERAL REMARKS: Description and illustration of alate adult male by Koteja & Azar (2008).

SYSTEMATICS: This species was described from alate adult male inclusion in Lower Cretaceous Lebanese amber (Koteja & Azar, 2008).

CITATIONS: HodgsoHa2013 [phylogeny, taxonomy: 799]; KotejaAz2008 [taxonomy, description, illustration, distribution: 159-161].