Family Coelostomidiidae


Coelostomidia Cockerell

NOMENCLATURE:

Coelostoma Maskell, 1880: 294. Type species: Coelostoma zealandica Maskell, by monotypy. Homonym of Coelostoma Brulle, 1835, in the Coleoptera; discovered by Cockerell, 1900a: 367.

Caelostoma; Maskell, 1884: 141. Misspelling of genus name. Notes: Mis-spelling of Coelostoma.

Coelostomidia Cockerell, 1900a: 367. Replacement name for Coelostoma Maskell, 1880.

Coelostomidea; MacGil1ivray, 1921: 86. Misspelling of genus name.

GENERAL REMARKS: Definition and characters by Morrison & Morrison (1922), Morrison (1928) and by Morales (1991).

SYSTEMATICS: The adult female of this genus has a well-developed, 11-segmented antennae, whereas in related genera (Ultracoelostoma, Paracoelostoma, Mimosicerya and Cryptokermes the antennae are reduced to an elongate cone (Morrison, 1928).

KEYS: Morales 1991: 24-25, 28-30 (female, male, nymph) [New Zealand]; Morrison 1928: 96-97 (female, larva) [Margarodidae].

CITATIONS: BenDov2005 [taxonomy, catalogue: 26]; Cocker1900a [taxonomy: 367]; Cocker1902q [taxonomy: 258]; Fernal1903b [catalogue: 30]; Foldi2001a [taxonomy: 198]; GullanCo2007 [taxonomy: 413-425]; GullanSj2001 [taxonomy: 257-278]; HodgsoFo2006 [taxonomy: 174]; Lindin1937 [taxonomy: 181, 182]; MacGil1921 [taxonomy, description: 86]; Maskel1880 [taxonomy, description: 294]; Maskel1884 [taxonomy: 141]; Maskel1887a [taxonomy: 90]; Maskel1890a [taxonomy: 280]; Maskel1893b [taxonomy: 241]; Morale1991 [taxonomy, description: 27-30]; Morris1927 [taxonomy: 102]; Morris1928 [taxonomy, description: 35,84-89,112-117,222]; MorrisMo1922 [taxonomy, description: 7-11]; MorrisMo1923 [taxonomy: 38]; MorrisMo1966 [taxonomy, catalogue: 25-26, 43].



Coelostomidia deboerae Morales

NOMENCLATURE:

Coelostomidia totarae Butcher, 1987: 146. Nomen nudum; discovered by Morales, 1991: 32.

Coelostomidia deboerae Morales, 1991: 30. Type data: NEW ZEALAND: South Island, Nelson, Maitai Vly, on Podocarpus totara; collected 17.i.1968, J.A. deBoer. Holotype female. Type depository: Auckland: New Zealand Arthropod Collection, Landcare Research, New Zealand; type no. 338. Described: female and first instar. Illust.



HOSTS: Epacridaceae: Dracophyllum [Butche1987]. Podocarpaceae: Dacrycarpus dacrydioides [Morale1991], Podocarpus taxifolia [Morale1991], Podocarpus totara [Morale1991].

DISTRIBUTION: Australasian: New Zealand (North Island [Morale1991], South Island [Morale1991]).

GENERAL REMARKS: Description and illustration of nymphs and adult female by Morales (1991).

KEYS: Morales 1991: 28-30 (female, male, nymph) [New Zealand].

CITATIONS: BenDov2005a [taxonomy, host, distribution, catalogue: 27]; Butche1987 [taxonomy: 146]; Foldi2001a [taxonomy: 198]; Morale1991 [taxonomy, description, illustration, host, distribution: 30-32, 74-76].



Coelostomidia jenniferae Morales

NOMENCLATURE:

Coelostomidia jenniferae Morales, 1991: 32. Type data: NEW ZEALAND: South Island, Southland, near Curio Bay, on Leptospermum sp., Feb. 1983, J.M. Cox. Holotype female. Type depository: Auckland: New Zealand Arthropod Collection, Landcare Research, New Zealand; type no. 83.2631. Described: female and first instar. Illust.



HOSTS: Araliaceae: Schefflera digitata [Morale1991]. Asteraceae: Olearia avicenniaefolia [Morale1991], Olearia colensoi [Morale1991], Olearia ilicifolia [Morale1991], Olearia traversii [Morale1991], Traversia baccharoides [Morale1991]. Cunoniaceae: Weinmannia racemosa [Morale1991]. Cupressaceae: Libocedrus bidwilli [Morale1991]. Elaeocarpaceae: Aristotelia [Morale1991]. Fagaceae: Nothofagus fusca [Morale1991], Nothofagus menziesii [Morale1991], Nothofagus solandri [Morale1991]. Myrtaceae: Leptospermum [Morale1991], Leptospermum scoparium [Morale1991], Metrosideros [Morale1991]. Phyllocladaceae: Phyllocladus alpinus [Morale1991]. Podocarpaceae: Prumnopitys taxifolia [Morale1991]. Winteraceae: Pseudowintera colorata [Morale1991].

DISTRIBUTION: Australasian: New Zealand (South Island [Morale1991]).

GENERAL REMARKS: Description and illustration of adult female by Morales (1991).

KEYS: Morales 1991: 28-30 (female, male, nymph) [New Zealand].

CITATIONS: BenDov2005a [taxonomy, host, distribution, catalogue: 27]; Foldi2001a [taxonomy: 108]; Morale1991 [taxonomy, description, illustration, host, distribution: 32-34, 77-79].



Coelostomidia montana Green

NOMENCLATURE:

Coelostomidia montana Green, 1929: 370. Type data: NEW ZEALAND: South Island, Arthur's Pass, Altitude 3300 ft., on Dracophyllum traversii, i.1923. Lectotype female, by subsequent designation Morales, 1991: 34. Type depository: London: The Natural History Museum, England, UK. Described: female. Illust.



HOSTS: Myrtaceae: Leptospermum ericoides [Morale1991]. Podocarpaceae: Dracophyllum [Morale1991], Dracophyllum traversii [Green1929, Morale1991].

DISTRIBUTION: Australasian: New Zealand (South Island [Morale1991]).

GENERAL REMARKS: Description and illustration of adult female and nymphs by Morales (1991).

KEYS: Hodgson & Foldi 2006: 191-192 (male) [Males of Coelostomidiidae].

CITATIONS: BenDov2005a [taxonomy, host, distribution, catalogue: 28]; Butche1987 [host, distribution: 146]; Foldi2001a [taxonomy: 198]; Green1929 [taxonomy, description, illustration, host, distribution: 370-372]; HodgsoFo2006 [taxonomy: 191]; Morale1991 [taxonomy, description, illustration, host, distribution: 34-36, 80-85].



Coelostomidia pilosa (Maskell)

NOMENCLATURE:

Coelostoma pilosum Maskell, 1891: 29. Type data: NEW ZEALAND: Reefton district, on various trees, Podocarpus totara and Fagus spp., in forests. Lectotype female, by subsequent designation Morales, 1991: 38. Type depository: Auckland: New Zealand Arthropod Collection, Landcare Research, New Zealand. Described: female. Illust.

Coelostomidia pilosa; Cockerell, 1902q: 258. Change of combination.

Coelostomidia pilosam; Morrison & Morrison, 1922: 10. Misspelling of species name.

COMMON NAME: shaggy giant scale [Morale1991].



HOSTS: Araliaceae: Neopanax [Morale1991]. Cunoniaceae: Weinmannia racemosa [Morale1991]. Cupressaceae: Libocedrus bidwilli [Morale1991]. Fagaceae: Fagus [Maskel1891], Nothofagus [Morale1991], Nothofagus solandri [Morale1991]. Liliaceae: Hebe [Morale1991]. Myrtaceae: Metrosideros [Morale1991]. Pandanaceae: Freycinetia [Morale1991]. Phyllocladaceae: Phyllocladus alpinus [Morale1991]. Poaceae [Morale1991]. Podocarpaceae: Dacrydium [Morale1991], Dacrydium biforme [Morale1991], Dacrydium cupressinum [Morale1991], Podocarpus totara [Maskel1891, Morale1991], Prumnopytis ferruginea [Morale1991], Prumnopytis taxifolia [Morale1991]. Rosaceae: Ripogonum scandens [Morale1991], Rubus australis [Morale1991]. Winteraceae: Pseudowintera colorata [Morale1991].

DISTRIBUTION: Australasian: New Zealand (North Island [Maskel1891, Morale1991], South Island [Morale1991]).

GENERAL REMARKS: Description and illustration of adult female and female nymph by Morrison & Morrison (1923) and by Morales (1991). Description and illustration of adult male by Hodgson & Foldi (2006).

STRUCTURE: Colour illustration of the general appearance of the adult female and adult male by Green (1922). Adult female dark-red in colour. At first naked; then with a thin covering of white meal, which gradually becomes thicker and more solid, until it assumes the appearance of a hard, granular shell, looking like a coating of lime; at gestation this increases to a snow-white mass of closely-felted cotton covering the insect and the eggs. Length variable, some specimens 0.5 inch (Maskell, 1891). Adult male darkish-red or brown in colour, but covered with hairs and with a good deal of whitish meal on the body and legs; length about 0.1 inch (Maskell, 1892).

KEYS: Hodgson & Foldi 2006: 191-192 (male) [Males of Coelostomidiidae]; Morrison 1928: 117 (female, male) [Species of Coelostomidia].

CITATIONS: BenDov2005a [taxonomy, host, distribution, catalogue: 28-29]; Cocker1902q [taxonomy: 258]; DeitzTo1980 [taxonomy: 51]; Fernal1903b [catalogue: 30]; Foldi2001a [taxonomy: 198]; Green1929 [distribution: 369]; GullanCo2007 [taxonomy: 413-425]; HodgsoFo2006 [taxonomy, description, illustration, host, distribution: 174-180,191]; HodgsoHa2013 [phylogeny, taxonomy: 796,799]; Maskel1891 [taxonomy, description, illustration, host, distribution: 29-30]; Maskel1892 [taxonomy, description, illustration: 49]; Morale1991 [taxonomy, description, illustration, host, distribution: 36-39, 86-90]; Morris1928 [taxonomy: 117, 225]; MorrisMo1922 [taxonomy: 10]; MorrisMo1923 [taxonomy, description, illustration, host, distribution: 38-40]; Vayssi1926 [taxonomy, host, distribution: 351].



Coelostomidia wairoensis (Maskell)

NOMENCLATURE:

Caelostoma wairoense Maskell, 1884: 141. Type data: NEW ZEALAND: District of Wairoa, on Leptospermum sp.; collected June 1883, by Mr. T. Cheeseman. Lectotype male, by subsequent designation Morales, 1991: 39. Type depository: Auckland: New Zealand Arthropod Collection, Landcare Research, New Zealand. Described: male. Illust.

Caelostoma wairoensis; Maskell, 1884: 141. Misspelling of genus name.

Coelostomidia wairoensis; Cockerell, 1902q: 258. Change of combination.

COMMON NAME: manuka giant scale [Morale1991].



HOSTS: Agavaceae: Phormium [Maskel1884]. Myrtaceae: Kunzea ericoides [Morale1991], Leptospermum [Maskel1884, Butche1987], Leptospermum ericoides [Green1929, Morale1991], Leptospermum scoparium [Morale1991], Metrosideros [Morale1991].

DISTRIBUTION: Australasian: New Zealand [Maskel1884].

GENERAL REMARKS: The original description by Maskell (1884) was based on the adult male only. Description and illustration of adult male and nymphs by Morales (1991). Description and illustration of adult male by Hodgson & Foldi (2006).

KEYS: Hodgson & Foldi 2006: 191-192 (male) [Males of Coelostomidiidae].

CITATIONS: BenDov2005a [taxonomy, host, distribution, catalogue: 29]; Butche1987 [host, distribution: 146]; Cocker1902q [taxonomy, distribution: 258]; DeitzTo1980 [taxonomy: 51]; Fernal1903b [catalogue: 31]; Foldi2001a [taxonomy: 198]; GardneBe2008 [life history, ecology, host, distribution: 195-203]; Green1929 [taxonomy, description, illustration, host, distribution: 369-370]; GullanCo2007 [taxonomy: 413-425]; HodgsoFo2006 [taxonomy, description, illustration, host, distribution: 180-185]; HodgsoHa2013 [phylogeny, taxonomy: 796,799]; Maskel1884 [taxonomy, description, illustration, host, distribution: 141-143]; Maskel1887a [taxonomy, description, host, distribution, economic importance: 109]; Morale1991 [taxonomy, description, illustration, host, distribution: 39-41, 91-05]; Morris1928 [taxonomy: 117, 227]; MorrisMo1922 [taxonomy: 10]; MorrisMo1923 [taxonomy: 41].



Coelostomidia zealandica (Maskell)

NOMENCLATURE:

Coelostoma zealandica Maskell, 1880: 294. Type data: NEW ZEALAND: Otago on bark of large trees, and Canterbury buried in the ground and in the chinks of rocks; W.M. Maskell, April 1879. Lectotype larva, by subsequent designation Morales, 1991: 43. Type depository: Auckland: New Zealand Arthropod Collection, Landcare Research, New Zealand. Described: larva. Illust.

Caelostoma zealandicum; Maskell, 1884: 141. Misspelling of genus name.

Coelostomidia zealandica; Cockerell, 1900: 367. Change of combination.

COMMON NAME: great giant scale [Morale1991].



FOE: Pteromalidae: Proshizonotus [Noyes2004].

HOSTS: Araliaceae: Pseudopanax [Morale1991]. Asteraceae: Cassinia [Morale1991]. Elaeocarpaceae: Aristotelia serrata [Morale1991]. Epacridaceae: Cyathodes colensoi [Morale1991], Cyathodes fraseri [Morale1991]. Fagaceae: Nothofagus [Morale1991], Nothofagus menziesii [Morale1991]. Griseliniaceae: Griselinia [Morale1991]. Liliaceae: Collospermum [Morale1991]. Loranthaceae: Tupeia antarctica [Morale1991]. Malvaceae: Plagianthus [Morale1991]. Myoporaceae: Myoporum laetum [Morale1991]. Pittosporaceae: Pittosporum tenuifolium [Morale1991]. Podocarpaceae: Podocarpus totara [Morale1991]. Polygonaceae: Muehlenbeckia australis [Green1929, Morale1991], Muehlenbeckia complexa [Morale1991]. Rhamnaceae: Discaria [Morale1991]. Rosaceae: Rubus australis [Morale1991], Rubus schmideloides [Morale1991]. Scrophulariaceae: Hebe [Morale1991]. Thymelaeaceae: Pimelea [Morale1991].

DISTRIBUTION: Australasian: New Zealand (North Island [Morale1991], South Island [Morale1991]).

GENERAL REMARKS: Description and illustration of nymph, adult female and adult male by Morrison & Morrison (1922) and by Morrison (1928). Description and illustration of adult female and nymphs given by Morales (1991).

STRUCTURE: Maskell (1880) described the female as: "The adult female is brick-red in colour, reaching 0.5 inch in length and more than 0.25 inch in width. It is fat, corrugated, slug-like; there are eleven or twelve corrugations, those toward the head being the widest. It is surrounded by a thick envelope of white cotton, in which it deposits the eggs, which are oval and brick-red in colour".

KEYS: Hodgson & Foldi 2006: 191-192 (male) [Males of Coelostomidiidae]; Morrison 1928: 117 (female, male) [Species of Coelostomidia].

CITATIONS: BenDov2005a [taxonomy, host, distribution, catalogue: 29-30]; Cocker1900a [taxonomy: 367]; DeitzTo1980 [taxonomy: 51]; Fernal1903b [catalogue]; Foldi2001a [taxonomy: 198]; GardneBe2010 [host, distribution, ecology, life history: 295-303]; Green1929 [host, distribution: 369]; HodgsoFo2006 [taxonomy: 192]; HubbarPe1898 [taxonomy: 17]; Maskel1880 [taxonomy, description, illustration, host, distribution: 294-299]; Maskel1882 [taxonomy, description, illustration, host, distribution: 226-228]; Maskel1884 [taxonomy, description: 141]; Maskel1887 [taxonomy: 46]; Maskel1887a [taxonomy, host, distribution, economic importance: 107]; Maskel1889 [taxonomy: 101]; Morale1991 [taxonomy, description, illustration, host, distribution: 41-44, 96-99]; Morris1928 [taxonomy, description, illustration, host, distribution: 112-117, 227]; MorrisMo1922 [taxonomy, description, illustration, host, distribution: 7-11]; Noyes2004 [biological control]; Towns2003 [host, distribution, life history, ecology: 329-335]; VeitchCl2003 [host, distribution, life history, ecology].



Ultracoelostoma Cockerell

NOMENCLATURE:

Coelostomidia (Ultracoelostoma) Cockerell, 1902g: 114. Type species: Coelostoma assimile Maskell, by monotypy and original designation.

Ultracoelostoma; Morrison & Morrison, 1922: 11. Change of status.

GENERAL REMARKS: Definition and characters by Morrison (1928) and by Morales (1991).

SYSTEMATICS: Main characteristics (adult female) of this genus are: antennae reduced 5-7 segmented; legs reduced to stout cones, bearing a claw and digitules; labium absent; seven pairs of abdominal spiracles.

KEYS: Morales 1991: 24-25, 46 (female, male, nymph) [New Zealand]; Morrison 1928: 96-97 (female, larva) [Margarodidae].

CITATIONS: BenDov2005a [taxonomy, catalogue: 40-41]; Cocker1902q [taxonomy: 258]; Foldi2001a [taxonomy: 218]; GullanCo2007 [taxonomy: 413-425]; GullanSj2001 [taxonomy: 257-278]; HodgsoFo2006 [taxonomy: 185]; Lindin1937 [taxonomy: 197]; MacGil1921 [taxonomy, description: 87]; Morale1991 [taxonomy, description: 45-46]; Morris1928 [taxonomy, description: 13-16,23,96,108,220]; MorrisMo1922 [taxonomy, description: 11-14]; MorrisMo1966 [taxonomy, catalogue: 201].



Ultracoelostoma assimile (Maskell)

NOMENCLATURE:

Coelostoma assimile Maskell, 1890: 153. Type data: NEW ZEALAND: Reefton district, on Fagus sp., April 1890, W.M. Maskell. Lectotype female, by subsequent designation Morales, 1991: 47. Type depository: Auckland: New Zealand Arthropod Collection, Landcare Research, New Zealand. Described: female. Illust.

Coelostomidia assimilis; Cockerell, 1902q: 258. Change of combination.

Coelostomidia (Ultracoelostoma) assimile; Cockerell, 1902q: 258. Change of combination.

Ultracoelostoma assimilis; MacGillivray, 1921: 87. Change of combination.

Ultracoelostoma assimile; Morrison & Morrison, 1922: 11. Change of combination.

COMMON NAMES: sooty beech scale [MoraleHiWa1988].



FOES: COLEOPTERA Cleridae: Metaxina ornata Broun [MoraleHiWa1988]. Melandryiidae: Doxozilora punctata Broun [MoraleHiWa1988].

HOSTS: Cunoniaceae: Weinmannia [Butche1987]. Fagaceae: Fagus [Maskel1890], Fagus fusca [Maskel1891], Fagus menziesii [Maskel1891]. Monimiaceae: Laurelia [Butche1987]. Phyllocladaceae: Phyllocladus trichomanoides [Maskel1891].

DISTRIBUTION: Australasian: New Zealand (North Island [Maskel1890, Butche1987, Morale1991], South Island [Morale1991]).

GENERAL REMARKS: Description and illustration of nymph, adult female and adult male by Morrison & Morrison (1922) and by Morrison (1928). Description and illustration of adult female and nymphs by Morales (1991).

STRUCTURE: Adult female reddish-brown, occupying a deep pit in the twig it lives on (mostly in the axils), the mouth of the pit being covered with the mass of thick yellow wax, of irregularly globular shape, already formed by the second-stage female. The cavity burrowed out is frequently quite deep in the bark, and even seems sometimes to extend into the wool. The dimensions very: some of the waxy coverings are only about 1/15 inch, others reach 1/8 inch. The wax is very hard (Maskell, 1891).

ECONOMIC IMPORTANCE AND CONTROL: Regarded as the most important species, among the honeydew secreting Hompterans, for apiculture in New Zealand, (Walton, 1979; Crozier, 1981; Gaze & Clout, 1983; Airborne Honey, New Zealand, 2003). The annual yield of honeydew honey at Canterbury plains foothill forest (South Island, New Zealand) was estimated at 3302 tones (Walton, 1979).

KEYS: Hodgson & Foldi 2006: 191-192 (male) [Males of Coelostomidiidae].

CITATIONS: AirborHoLt1999 [host, distribution, economic importance, chemistry]; AstwooLeMa1998 [distribution, chemistry: 4958-4962]; BallAr2007 [taxonomy, molecular biology, molecular data: 1-14]; BeckhLu1998 [distribution, chemistry: 149-152]; Belton1979 [host, distribution, economic importance: 4-6]; BenDov2005a [taxonomy, host, distribution, catalogue, economic importance: 41-42]; Butche1987 [host, distribution, life history, biological control: 143-147]; Cocker1902q [taxonomy, distribution: 258]; Crozie1981 [host, distribution, economic importance: 200-209]; DeitzTo1980 [taxonomy: 50]; Dumble1967 [host, distribution, life history: 39-40]; DunganTuKe2007 [life history, ecology, physiology, host, distribution: 603-613]; Fernal1903b [catalogue: 30]; Foldi2001a [taxonomy: 218]; GazeCl1983 [economic importance, honeydew, host, distribution: 33-37]; Green1929 [host, distribution: 369]; GullanCo2007 [taxonomy: 413-425]; HodgsoFo2006 [taxonomy, description, illustration, host, distribution: 186-191]; HubbarPe1898 [taxonomy: 17]; Kelly1990 [host, distribution, life history, physiology, economic importance: 53-58]; MacGil1921 [taxonomy, host, distribution: 87]; Maskel1890 [taxonomy, description, illustration, host, distribution: 153-154]; Maskel1891 [taxonomy, description, illustration, host, distribution: 30-33]; Morale1991 [taxonomy, description, illustration, host, distribution: 46-49,100-102]; MoraleHiWa1988 [life history, host, distribution, biological control, economic importance: 24-37]; Morris1928 [taxonomy, description, illustration, host, distribution: 108-112,222]; MorrisMo1922 [taxonomy, description, illustration, host, distribution: 11-14]; Vayssi1926 [taxonomy, host, distribution: 351]; Walton1979 [economic importance, life history, host, distribution: 6-9]; WardhaBlGr2006 [host, distribution, life history, ecology: 185-195]; WardhaDi2005 [host, distribution, life history, ecology: 733-738]; WardhaDi2006a [host, distribution, life history, ecology: 1-9].



Ultracoelostoma brittini Morales

NOMENCLATURE:

Ultracoelostoma brittini Morales, 1991: 49. Type data: NEW ZEALAND: South Island, Maruia, on Nothofagus menziesii; collected 18.12.1935 by G. Brittin. Holotype female. Type depository: Auckland: New Zealand Arthropod Collection, Landcare Research, New Zealand. Described: female. Illust.



HOSTS: Fagaceae: Nothofagus fusca [Morale1991], Nothofagus menziesii [Morale1991], Nothofagus solandri [Morale1991].

DISTRIBUTION: Australasian: New Zealand (South Island [Morale1991]).

GENERAL REMARKS: Description and illustration of female, male and nymphs by Morales (1991).

ECONOMIC IMPORTANCE AND CONTROL: The honeydew secretion of this species, in the south island of New Zealand, is regarded an important food source for honeybees, Apis mellifera, and for the wasps Vespula germanica and V. vulgaris (Markwell et al., 1993; Airborne Honey, 2003).

CITATIONS: AirborHoLt1999 [host, distribution, economic importance, chemistry]; BallAr2007 [taxonomy, molecular biology, molecular data: 1-14]; BeggsKaWa2005 [host, distribution, biological control: 105-115]; BenDov2005a [taxonomy, host, distribution, catalogue: 42]; DunganBeWa2004 [host, distribution, economic importance, life history, chemistry: 283-288]; Foldi2001a [taxonomy: 218]; KellyStHu1992 [host, distribution, life history, physiology, economic importance: 69-75]; MarkweKeDu1993 [host, distribution, economic importance, life history: 85-93]; Morale1991 [taxonomy, description, illustration, host, distribution: 49-51, 103-109]; WardhaDi2006a [host, distribution, life history, ecology: 1-9].



Ultracoelostoma dracophylli Morales

NOMENCLATURE:

Ultracoelostoma dracophylli Butcher, 1987: 146. Nomen nudum; discovered by Morales, 1991: 51.

Ultracoelostoma dracophylli Morales, 1991: 51. Type data: NEW ZEALAND: South Island, North Canterbury, Arthur's Pass, on Dracophyllum longifolium, 23.1.1983, C.F. Butcher. Holotype female. Type depository: Auckland: New Zealand Arthropod Collection, Landcare Research, New Zealand; type no. 83.284a. Described: female. Illust.



HOSTS: Epacridaceae: Archeria [Morale1991], Dracophyllum [Butche1987], Dracophyllum filiforme [Morale1991], Dracophyllum longifolium [Morale1991], Dracophyllum lyalli [Morale1991], Dracophyllum paludosum [Morale1991].

DISTRIBUTION: Australasian: New Zealand (South Island [Morale1991]).

GENERAL REMARKS: Description and illustration of adult female and nymphs by Morales (1991).

CITATIONS: BenDov2005a [taxonomy, host, distribution, catalogue: 42-43]; Butche1987 [taxonomy: 146]; Foldi2001a [taxonomy: 218]; Morale1991 [taxonomy, description, illustration, host, distribution: 51-53, 110-113].





Genera Removed from Family


Cryptokermes

No valid record found for this genus

NOMENCLATURE:

Cryptokermes Hempel, 1900: 398. Notes: Cryptokermes has been transferred to the Monophlebidae.

Cryptokermes Hempel, 1900a: 380,398. Notes: Current status: Cryptokermes Hempel in Monophlebidae.



Mimosicerya

No valid record found for this genus

NOMENCLATURE:

Icerya (Mimosicerya) Cockerell, 1902q: 233. Notes: Current status: Mimosicerya (Cockerell) in Mimo.



Neocoelostoma

No valid record found for this genus

NOMENCLATURE:

Neocoelostoma Hempel, 1932: 310. Notes: Current status: Neocoelostoma Hempel in Monophlebidae.



Paracoelostoma

No valid record found for this genus

NOMENCLATURE:

Paracoelostoma Morrison, 1927: 102. Notes: Current status: Paracoelostoma Morrison in Monophlebidae.



Species Removed from Family


Cryptokermes brasiliensis

No valid record found for this species

NOMENCLATURE:

Cryptokermes brasiliensis Hempel, 1900a: 398. Notes: Current status: Cryptokermes brasiliensis Hempel in Monophlebidae.

Cryptokermes mexicanus

No valid record found for this species

NOMENCLATURE:

Cryptokermes mexicanus Morrison, 1927: 103. Notes: Current status: Cryptokermes mexicanus Morrison in Monophlebidae.

Cryptokermes mimosae

No valid record found for this species

NOMENCLATURE:

Cryptokermes mimosae Foldi, 1995a: 173. Notes: Current status: Cryptokermes mimosae Foldi in Monophlebidae.

Cryptokermes oaxacaensis

No valid record found for this species

NOMENCLATURE:

Cryptokermes oaxacaensis Foldi, 2011: 484-487. Notes: Current status: Cryptokermes oaxacaensis Foldi in Monophlebidae.

Mimosicerya hempeli

No valid record found for this species

NOMENCLATURE:

Icerya (Crypticerya) hempeli Cockerell, 1899c: 43. Notes: Current status: Mimosicerya hempeli (Cockerell) in Monophlebidae.

Mimosicerya williamsi

No valid record found for this species

NOMENCLATURE:

Mimosicerya williamsi Foldi, 2009: 28. Notes: Current status: Mimosicerya williamsi Foldi in Monophlebidae.

Neocoelostoma xerophila

No valid record found for this species

NOMENCLATURE:

Neocoelostoma xerophila Hempel, 1932: 311. Notes: Current status: Neocoelostoma xerophila Hempel in Monophlebidae.

Paracoelostoma peruvianum

No valid record found for this species

NOMENCLATURE:

Paracoelostoma peruviana Morrison, 1927: 103. Notes: Current status: Paracoelostoma peruvianum Morrison in Monophlebidae.