Vasseur, R., & Bénassy, C. 1953 [On the parasites of the San José scale (Quadraspidiotus perniciosus Comst.) in the Lyon region and the effect of the climatic conditions.]. Annales des Epiphyties 4: 283-290.
Notes: Original title: [Sur la faune parasitaire du pou de San José (Quadraspidiotus perniciosus Comst.) dans la région lyonnaise et ses relations avec les conditions climatiques.] The two most effective parasites of Q. perniciosus in the Lyon region are Aphytis proclia and A. mytilaspidis. The cold temperatures of winter diminish their effectiveness.