Takagi, S., & Hodgson, C.J. 2005 A new dipterocarp-associated gall-inhabiting coccoid from Borneo (Homoptera: Coccoidea).. Insecta Matsumurana (New Series) 61: 11-41.

Notes: Danumococcus parashoreae, gen. et sp. nov., is described on the basis of specimens collected at Danum Valley, Sabah, Malaysia, on the leaves of Parashorea tomentella (Dipterocarpaceae). The adult female inhabits a small lenticular gall induced on the blade. The gall has a small round opening at the centre of the roof. The male normally occurs outside the gall, and secretes a loose mass of wax to form a cocoon around the body in the second instar. The adult female is extremely degenerate, having no setiferous antennae, no legs, and no disc pores. It has been unusually modified in several other features and is provided with some peculiar, remarkable developed structures, apparently associated with the opening in the roof of the lenticular gall. The first-instar female and the first- and second-instar and adult males, the stages probably not or less modified in association with the gall structure, have some features similar to those in Beesonia, Magalorea, Gallacoccus, and Echinogalla. The view is adopted tentatively that the gall-inhabiting coccoids of these genera and Danumococcus, though far from uniform, belong to the same phylogenetic group, the family Beesoniidae, which has evolved mainly on dipterocarps. However, the second-instar male of Danumococcus is provided with clusters of deeply concave disc pores similar to those on Dactylopius. This fact may suggest the presence of some phylogenetic relationship between Danumococcus and Dactylopius and, further, may promote the concept of a broader natural group comprising all the genera mentioned above and others now in separate families. A cladistic analysis based on macropterous male characters shows that D. parashoreae belongs to the Beesoniidae and also that the Beesoniidae and the Stictococcidae fall within a larger clade which includes the Dactylopiidae and some gall-inducing eriococcids.