Steyn, J.J. 1954 The effect of the cosmopolitan brown house ant (Pheidole megacephala F.) on citrus red scale (Aonidiella aurantii Mask.) at Letaba.. Journal of the Entomological Society of Southern Africa 17: 252-264.

Notes: P. megacephala was found to be the cause of epidemics of A. aurantii on citrus; noctrnal feeding periodicity of this ant was suggested as the reason why this correlation had not previously been revealed.