Shcherbakov, D.E. 2007 Extinct four-winged precoccids and the ancestry of scale insects and aphids (Hemiptera).. Russian Entomological Journal 16(1): 47-62.

Notes: ABSTRACT. The Naibiidae fam.n. (Triassic-Paleocene) is described, comprising Coccavus supercubitus gen. et sp.n. (Coccavinae subfam.n.), Panirena sukatshevae gen. et sp.n., P. tenella sp.n. and Naibia zherichini gen. et sp.n. This family, demonstrating several coccid apomorphies combined with still aphid-like habitus, is placed in the infraorder Coccomorpha s.l. (as a separate superfamily) and considered to be ancestral to the true scale insects. Close relationship of Aphidomorpha and Coccomorpha is confirmed by fossils, and both groups are derivable from the same stem lineage represented by Boreoscytidae (Permian), Pincombeidae (Permian-Triassic) and Simulaphididae fam.n. (Permian Simulaphis shaposhnikovi gen. et sp.n., possibly also Triassic). These three families are included in the suborder Aphidinea s.l. (as infraorder Pincombeomorpha) along with aphids and coccids. New synonymies are established and Dinoscyta microcephala gen. et sp.n. is described in Boreoscytidae. All the taxa dealt with are keyed, and a phylogram of Aphidinea is proposed. Evolution of coccid forewing is discussed. KEY WORDS: scale insects, coccids, aphids, Coccomorpha, Aphidomorpha, Sternorrhyncha, Hemiptera, Boreoscytidae, Pincombeidae, Simulaphididae, Naibiidae, Ortheziidae, fossils, Sakhalin amber, phylogeny, Permian, Triassic, Jurassic, Paleocene.