Riom, J., & Fabre, J.P. 1977 Étude biologique et écologique de la cochenille du Pin maritime, Matsucoccus feytaudi Ducasse, 1942 (Coccoidea, Margarodidae, Xylococcinae) sans le Sud-Est de la France. II. Régulation du cycle annuel, comportements des stades mobiles.. Annales de Zoologie - Ecologie Animale 9: 181-209.

Notes: Title in English: [Biological and ecological study of the maritime pine scale Matsucoccus feytaudi Ducasse, 1942 (Coccoidea, Margarodidae, Xylococcinae) in south-eastern France. II. - Regulation of the annual cycle, behaviour of the mobile stages.] Further details are given of field and laboratory studies in France on the biology of Matsucoccus feytaudi Ducasse, which is injurious to maritime pine [Pinus pinaster] in the south of the country. Development of the scale insect is limited to 1 generation a year because of the time taken to reach the first nymphal moult; the first instar lasted 100-200 days (about 140 days in the field). High temperatures in July and August induced aestivation at the end of the instar, and temperatures of 10-15 deg C were necessary to induce the moult. Studies on dispersal within the tree showed that the first-instar nymphs remained immobile for 12-15 days after hatching at 15 deg C and 3-4 days after hatching at 25 deg C; 75% of the population was then mobile for 8-43 h, showing positive phototropism and negative geotropism. About 50% of the population covered 25-45 cm, according to the population density; only 0.3% covered distances of 2.5-3 m. Population density affected the distance covered, but not thedirection of the movements. Pronymphs and adult females tended to move down towards the base of the tree, and adult males showed a tendency to move upwards.