Hughes-Schrader, S. 1963 Hermaphroditism in an African coccid, with notes on other Margarodids (Coccoidea-Homoptera).. Journal of Morphology 113: 173-184.

Notes: Detailed study on cytology of Icerya tremae, maxima, aegyptiaca, bimaculata and Kuwania oligostigma; gives chromosome complement for each sp.; discusses cytotaxonomy.