Hadzibejli, Z.K., & Gelovani, L.V. 1983 [The importance of intraspecific variability for the determination of scale insects.]. Fauna i Ecologiya Bespozv. Zhivontn. Gruzii 178-195.

Notes: Studies 4 coccoid species; discusses differences in size, shape and color due to feeding locations on different plants; seasonal changes in males and females.