Foldi, I. 2005 The Matsucoccidae in the Mediterranean basin with a world list of species (Hemiptera: Sternorrhyncha: Coccoidea).. Annales de la Société Entomologique de France 40(2)(2004): 145-168.


  • description
  • illustration
  • list
  • taxonomy
  • Notes: The matsucoccoid scale insects inhabit the northern hemisphere, feed exclusively on Pinus spp., and some are very serious pests. This taxonomic revision of the Mediterranean species supports their status as valid species, despite the wide variation of certain morphological characters caused by environmental factors. For the genus redescription and illustrations, Matsucoccus matsumurae Kuwana, the type species of the genus from Japan is used. The Mediterranean species, M. feytaudi, M. pini and M. josephi are redescribed and illustrated. A key for their identification is provided using morphological characters and quantitative features. In M. feytaudi, all stages of development of both the male and female are described and illustrated. Lectotypes of M. feytaudi and M. pini are designated. Some information on the biology, host plants, economic importance, natural enemies and distribution of all species is given. A world list of extant and fossil species of Matsucoccidae is provided along with some pertinent (host plant, feeding site, distribution) information.