Valid Names Results

Bessenayla balachowskyi is now Fonscolombia tomlinii (Newstead, 1892) (Pseudococcidae: Fonscolombia)

Nomenclatural History

Ripersia tomlinii Newstead 1892: 146. Type data: CHANNEL ISLANDS: Guernsey, on grass roots in ants' nests. Syntypes, female, Type depository: London: The Natural History Museum, England, UK; accepted valid name
Ripersia montana Newstead 1898: 97. Type data: FRANCE: Haute Savoie, Argentieres, on roots of grass and composites. Lectotype, female, by subsequent designation (Matile2004,75). Type depository: London: The Natural History Museum, England, UK; junior synonym (discovered by Matile2004, 75).
Ripersia exul exul Green 1924b: 46. Type data: CHANNEL ISLANDS: Guernsey, in nests of ants. Syntypes, female, Type depository: London: The Natural History Museum, England, UK; junior synonym (discovered by Willia1962, 26).
Ripersia mesnili Balachowsky 1934a: 67. Type data: FRANCE: Corsica, Niolo, on Trisetum. Holotype, female, Type depository: Paris: Museum National d'Histoire naturelle, France; junior synonym (discovered by Willia1962, 26).
Pseudococcus bufo Kiritchenko 1936a: 149. Type data: UKRAINE: Odessa Oblast, Odessa, on roots of Gramineae, Krym, Kekeneiz, on roots of Poa bulbosa. Syntypes, female, Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; junior synonym (discovered by Danzig1971, 373).
Phenacoccus agropyri Borchsenius 1937: 52. Type data: UKRAINE: Northern Caucasus, Shuntk Station, in ditch near rail way station, on couch-grass roots; collected N.B. 22.v.1934.. Lectotype, female, by subsequent designation (Danzig2007,367). Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; junior synonym (discovered by Danzig1971, 373).
Ripersia exul allosiana Goux 1945: 37. Type data: FRANCE: Lac d'Allos, Basses-Alpes, on the roots of grass. Lectotype, female, by subsequent designation (Matile2004,75). Type depository: Paris: Museum National d'Histoire naturelle, France; junior synonym (discovered by Matile2004, 75). Illustr.
Phenacoccopsis agropyri (Borchsenius, 1937); Borchsenius 1949: 200. change of combination
Phenacoccopsis bufo (Kiritchenko, 1936); Borchsenius 1949: 202. change of combination
Phenacoccopsis nassonovi Borchsenius 1949: 201. Type data: RUSSIA: St. Peterburg [= Leningrad] Oblast, on grass. Lectotype, female, by subsequent designation (Danzig2007,367). Type depository: St. Petersburg: Zoological Museum, Academy of Science, Russia; junior synonym (discovered by Danzig1971, 373).
Euripersia tomlinii (Newstead, 1892); Williams 1962: 26. change of combination
Euripersia montana (Newstead, 1898); Kozár & Walter 1985: 68. change of combination
Bessenayla balachowskyi Goux 1988: 6. Type data: FRANCE: Courzieu near Lynos, on Aira sp.. Holotype, female, Type depository: Paris: Museum National d'Histoire naturelle, France; junior synonym (discovered by DanzigGa2014, 241). Illustr.
Bessenayla balachoswkyi Goux, 1988; Goux 1988a: 6. misspelling of species epithet
Fonscolombia tomlinii (Newstead, 1892); Danzig 1994: 46. change of combination
Conicoccus montrottieri Goux 1994: 43. Type data: FRANCE: Rhone, Bessenay, on Gramineae; collected 12.viii.1952, by L. Goux. Lectotype, female, by subsequent designation (Matile2004,75). Type depository: Paris: Museum National d'Histoire naturelle, France; junior synonym (discovered by Matile2004, 75).
Bessenayla balachoswkyi Goux, 1988; Ben-Dov 1994: 61. misspelling of species epithet Notes: Bessenayla balachowskyi mis-spelled Bessenayla balachoswkyi.
Euripersia exul allosiana (Goux, 1945); Matile-Ferrero & Ben-Dov 1995: 258. change of combination
Eurypersia tomlini; Lagowska & Koteja 1996: 33. misspelling of genus name
Phenacoccopsis nasononi Borchsenius, 1949; Tao 1999: 14. misspelling of species epithet
Fonscolombia tomlini (Newstead, 1892); Gertsson 2011: 40. misspelling of species epithet

Common Names

Tomlin's mealybug KosztaKo1988F

Ecological Associates

Hosts:

Families: 6 | Genera: 26

Apiaceae

Daucus carota | BenDov1994

Asteraceae

Artemisia dracunculus | BenDov1994 Danzig1974
Asteraceae | BenDov1994 Marott1987a Newste1898
Tragopogon | BenDov1994 BorchsTe1956

Crassulaceae

Sedum | Danzig1994

Cyperaceae

Carex | BenDov1994 Danzig1959

Poaceae

Agrostis capillaris | BatsanKaKi2017
Aira | BenDov1994 Goux1988a
Arrhenatherum elatius | BenDov1994 Schmut1952
Bromus | BenDov1994 KosztaKo1988F
Corynephorus canescens | BenDov1994 Schmut1952
Dactylis glomerata | BenDov1994 Mateso1968
Deschampsia flexuosa | BenDov1994 Schmut1952
Elymus repens | BenDov1994 BorchsTe1956 TerGri1966 | (= Agropyron repens)
Festuca glauca | BenDov1994 Zahrad1987
Festuca ovina | BenDov1994 Koteja1972
Festuca rupicola | BenDov1994 Mateso1968 | (= Festuca sulcata)
Helictotrichon pubescens | BenDov1994 Mateso1968 | (= Avenastrum pubescens)
Hordeum | BenDov1994 TerGri1966
Koeleria macrantha | BenDov1994 Mateso1968 | (= Koeleria gracilis)
Leymus angustus | BenDov1994 Mateso1968 | (= Elymus angustus)
Phleum phleoides | BenDov1994 Mateso1968
Poa bulbosa | BenDov1994 Kiritc1936a
Poa compressa | BenDov1994 Koteja1972
Poa pratensis | BenDov1994 Mateso1968
Poa versicolor | BenDov1994 Mateso1968 | (= Poa stepposa)
Poaceae | BenDov1994 Goux1931 Goux1945 Marott1987a Newste1898
Psathyrostachys juncea | BenDov1994 Mateso1968 | (= Elymus junceus)
Puccinellia | BenDov1994 Mateso1968 | (= Atropis)
Stipa grandis | BenDov1994 Danzig1984
Stipa sibirica | BenDov1994 Danzig1984
Triticum | BenDov1994 TerGri1966
Triticum aestivum | BenDov1994 BorchsTe1956 | (= Triticum vulgare var. erythrospermum)
Triticum turgidum | BenDov1994 BorchsTe1956 | ssp. durum (= Triticum compactum erinaceum)

Rubiaceae

Galium | BenDov1994 KosztaKo1988F

Foes:

Families: 1 | Genera: 1

Encyrtidae

Ericydnus longicornis | KosztaKo1988F Schmut1955b

Geographic Distribution

Countries: 20

Armenia | BenDov1994 Borchs1949 Danzig2007 TerGri1966

China | BenDov1994 Tang1992 TangHaTa1992

Corsica | Foldi2003 Matile2004

Czech Republic | BenDov1994 Zahrad1987

Denmark | BenDov1994 KosztaKo1988F

France | BenDov1994 Foldi2001 Goux1988a | Balach1934a BenDov1994 Foldi2001 Goux1931 Goux1945 MatilePe2002 Newste1898

Georgia (Republic of) | BatsanKaKi2017

Germany | BenDov1994 Schmut1952 Schmut1980

Hungary | BenDov1994 KozarKiSa2004 KozarWa1986

Italy | BenDov1994 Leonar1920 LongoMaPe1995 Marott1987a Newste1900

Sicily | MazzeoRuLo1997 RussoMa1997

Kazakhstan | BenDov1994 Danzig2007 Mateso1968

Mongolia | BenDov1994 Danzig1980 Danzig1984a

Netherlands | BenDov1994 Reyne1951

Poland | BenDov1994 Koteja1974b KotejaZa1983 Lagows1996

Russia

Lipestsk Oblast | Gavril2003a
St. Petersburg (=Leningrad) Oblast | BenDov1994 Borchs1949
Yakutia-Sakha (=Yakut) AR | BenDov1994 Danzig1978a

Spain | Martin1985

Sweden | Gertss2000 Gertss2001

Switzerland | Danzig1994

Ukraine | BenDov1994 Borchs1949 Danzig2007

Krym (=Crimea) Oblast | BenDov1994 Borchs1937

United Kingdom

Channel Islands | BenDov1994 Newste1892
England | BenDov1994 Willia1962

Keys

Kaydan2014: pp.446 ( Adult (F) ) [Key to adult female Fonscolombia]
DanzigGa2014: pp.198-201 ( Adult (F) ) [Species of Fonscolombia]
Danzig2007: pp.364-365 ( Adult (F) ) [Fonscolombia species of Russia]
Matile2004: pp.76 ( Adult (F) ) [Fonscolombia species of France]
Tang1992: pp.447 ( Adult (F) ) [Palaearctic & Oriental regions]
KosztaKo1988F: pp.96 ( Adult (F) ) [Central Europe]
Terezn1975: pp.177 ( Adult (F) ) [Ukraine]
Borchs1949: pp.199 ( Adult (F) ) [Palaearctic region]

Remarks

Systematics: The new combination by Matile-Ferrero & Ben-Dov (1995) was based on characters of the species as given in Goux (1945).Goux (1988) page 8, published the species epithet mis-spelled Bessenayla balachoswkyi, but further in the original description it was correctly spelled balachowskyi.
Structure: Female up to 3.5 mm long, pink. Antennae 6-8-segmented. Claw denticle well developed. Anal apparatus complete; number of its pores and spinulae and length of setae vary in different populations. Circuli 2 or 3 in number, conves, sclerotized. Multilocular pores acattered on ventral surface of abdomen and metathorax. Quinquelocular pores scattered in medial zone of thoraxic sternites. Simple discoidal porees sparsely scallered on entire surface of body. Simple tubular ducts of one size forming trhnsverse rows on abdomen and randomly sparse on thorax. Cerarii numberint 2-3 pair. Small conical setae present on dorsum along with flagellate setae of different sizes. (Danzig & Gavrilov-Zimin, 2014)
Biology: Living on roots of the host plant.Occurring on the roots of a variety of herbaceous plants, especially grasses often found in association with ants. Life history, natural enemies and associated ant discussed by Kosztarab & Kozár (1988F).
General Remarks: Description and illustration of the adult female by Borchsenius (1939a), Williams (1962), Ter-Grigorian (1973) (as P. bufo), Tereznikova (1975), Zahradnik (1987), Kosztarab & Kozár (1988F), Tang (1992) and by Danzig (2007). Description of the adult female and first-instar nymph by Borchsenius (1949). Description and illustration of first-instar nymph by Ter-Grigorian (1973). Description of the adult female given by Leonardi (1920), Gomez-Menor Ortega (1937) and by Goux (1945).Good description and illustration of the adult female given by Goux (1988a).

Illustrations

Citations

Balach1934a: description, distribution, host, illustration, taxonomy, 67-71
BatsanKaKi2017: distribution, host, 263
BenDov1994: catalog, 61,62,156,157,474
Borchs1937: description, distribution, host, illustration, taxonomy, 52
Borchs1939a: description, distribution, host, illustration, taxonomy, 46-47
Borchs1949: description, distribution, host, taxonomy, 200-203
BorchsTe1956: distribution, host,
BugajNJuKa2021: distribution, host, list, 308
Cocker1897c: distribution, host, taxonomy, 240
Danzig1959: distribution, host, 445
Danzig1971: distribution, host, taxonomy, 373
Danzig1974: distribution, host, 69
Danzig1978a: distribution, host, 74
Danzig1980: distribution, host, 33
Danzig1984a: distribution, host, 33
Danzig1994: distribution, host, taxonomy, 46
Danzig2007: description, distribution, host, illustration, taxonomy, 367-368
DanzigGa2005: cytogenetics, taxonomy, 527-529
DanzigGa2014: description, distribution, host, illustration, taxonomy, 14,200,240-243
Foldi2001: distribution, 303-308
Foldi2003: distribution, host, 148
FoldiGe2018: distribution, list, 12
Gavril2003a: distribution, host, 109
Gavril2004: distribution, host, 526
Gavril2007: taxonomy, 1
Gavril2018: reproduction, 221
GavrilKu2005: distribution, taxonomy, 19-34
Gertss2000: distribution, host, 149
Gertss2001: distribution, 123-130
Gertss2011: distribution, 40
GomezM1937: description, distribution, host, taxonomy, 363-365
Goux1931: distribution, host, 332
Goux1945: description, distribution, host, taxonomy, 37
Goux1988a: description, distribution, host, illustration, taxonomy, 6-11
Goux1994: description, distribution, host, illustration, taxonomy, 41-43
Green1920: taxonomy, 122
Green1924b: description, distribution, host, illustration, taxonomy, 46-47
Jansen2001: distribution, host, 197-206
KalandSi2014: distribution,
Kaydan2014: taxonomy, 444,446
Kiritc1936a: description, distribution, host, illustration, taxonomy, 149-151
KosztaKo1988F: description, distribution, host, illustration, taxonomy, 97-98
Koteja1972: distribution, host, 567
Koteja1974b: distribution, taxonomy, 74
KotejaZa1983: distribution, host, 472
KozarKiSa2004: distribution, 57
KozarWa1985: catalog, taxonomy, 68
KozarWa1986: distribution, host, 115
LagowsGo2020: distribution, 70
LagowsKo1996: distribution, host, 31, 33
Leonar1920: description, distribution, host, taxonomy, 423
LongoMaPe1995: distribution, 118
LongoRu1990: distribution, host, 114
Marott1987a: distribution, host, 109
MarottTr1995a: taxonomy, 70-71
Martin1985: distribution, host, taxonomy, 98
Mateso1968: distribution, host, life history, 108-109
Matile2004: distribution, host, taxonomy, 75-76
MatileBe1995: taxonomy, 258
MatilePe2002: distribution, host, 351
Newste1892: description, distribution, host, illustration, taxonomy, 146-147
Newste1898: 97
Reyne1951: description, distribution, host, illustration, taxonomy, 189-206
RussoMa1997: distribution, 48
Schmut1952: distribution, host, 392
Schmut1955b: biological control, 515
SchmutHo2016: distribution, host, 23-24
SchmutHo2016: distribution, host, 33
SimonKa2011: distribution, 235
Tang1992: description, distribution, host, illustration, taxonomy, 455-456, 717
TangHaTa1992: distribution, host, 8
Tao1999: distribution, host, taxonomy, 14
TerGri1966: distribution, host, 87
TerGri1973: description, distribution, host, illustration, taxonomy, 125-127
Terezn1975: description, distribution, host, illustration, taxonomy, 180-182
Willia1962: description, distribution, host, illustration, taxonomy, 26-28
Willia2017a: catalog, list of species, 246, 248, 251
WilliaBe2015: taxonomy, 167
WilliaMa2012: distribution, 83,86
Zahrad1987: description, distribution, host, illustration, taxonomy, 355-357